1	"Abaxial calyx lobe fused"	"Coleus scutellarioides"
1	"Abaxial calyx lobes free"	2
2	"Stems, branches and the lower surface of the leaves with antrorse, non-glandular trichomes"	3
2	"Stems, branches and the lower surface of the leaves with divaricate to retrorse, non-glandular trichomes"	26
3	"Corolla with spots/speckles present at least on part of the upper lip (adaxial lobes and lateral lobes), and the upper lip approximately equal to or greater than the length of lower lip (abaxial lobe)"	4
3	"Corolla lacking spots/speckles, with upper lip approximately half the length of the lower lip"	7
4	"Leaf margin with up to six pairs of teeth"	5
4	"Leaf margin with 7–15 pairs of teeth"	6
5	"Succulent plant, leaf lamina broad-ovate to orbicular; corolla tube less than 3 times the width of calyx tube"	"Coleus verticillatus"
5	"Fleshy (but not succulent) plant, leaf lamina triangular ovate; corolla tube more than 3 times the width of calyx tube"	"Plectranthus saccatus"
6	"Stems and leaf adaxial surface with orange-coloured sessile glands; adaxial surface of leaf with veins strongly impressed; adaxial calyx lobe cordate to ovate; corolla tube swollen at mouth of calyx"	"Plectranthus ciliatus"
6	"Stems and leaf adaxial surface lacking orange-coloured sessile glands; adaxial surface of leaf flat with veins scarcely impressed; adaxial calyx lobe narrow-deltate; corolla tube attenuate without swelling at mouth of calyx"	"Plectranthus ecklonii"
7	"Leaf lamina serrate in upper half only"	"Coleus minutus"
7	"Leaf lamina serrate for entire length"	8
8	"Corolla > 25 mm long; flowering calyx with adaxial lobe apiculate, non-glandular trichomes on calyx tube at least twice the length of non-glandular trichomes on lobes (calyx bearded)"	"Coleus barbatus"
8	"Corolla < 20 mm long; flowering calyx with adaxial lobe acute to obtuse (not apiculate), non-glandular trichomes on calyx tube similar length to non-glandular trichomes on lobes (calyx not bearded)"	9
9	"Stems and leaves woolly comose"	"Coleus magnificus"
9	"Stems and leaves not woolly comose"	10
10	"Leaf lamina with 3–6 pairs of teeth"	11
10	"Leaf lamina with 6–23 pairs of teeth"	12
11	"Yellow-coloured sessile glands present on abaxial leaf surface and floral parts, seedling-derived stems with a fleshy tuberous base"	"Coleus arenicola"
11	"Coloured sessile glands absent on leaves and floral parts, seedling-derived stems without a fleshy tuberous base"	"Coleus ventosus"
12	"Leaf trichomes drying yellow; corolla 13–16 mm long; yellow-coloured sessile glands on leaves, branches, floral axis and calyces"	"Coleus blakei"
12	"Leaf trichomes uncoloured; corolla 7–13 mm; sessile glands yellow, orange or red on leaves, branches, floral axis and calyces if present"	13
13	"Stems with coloured sessile glands"	14
13	"Stems without coloured sessile glands (colourless sessile glands may be present)"	19
14	"Live foliage blue-silver; inflorescence axis without glandular trichomes"	"Coleus cyanophyllus"
14	"Live foliage green; inflorescence axis with glandular trichomes"	15
15	"Leaf lamina teeth 4–6 mm long; with yellow sessile glands"	"Coleus megadontus"
15	"Leaf lamina teeth up to c. 3 mm long; with orange/red sessile glands"	16
16	"Verticillasters 20–50-flowered; flowering calyx <1.8 mm long"	"Coleus congestus"
16	"Verticillasters 10–20-flowered; flowering calyx >1.8 mm long"	17
17	"Stems lacking to sometimes at most sparsely covered with coloured sessile glands, non-glandular trichomes to 1.8 mm long; leaf indumentum providing hoary appearance, especially when dry"	"Coleus gratus"
17	"Stems sparsely to densely covered with coloured sessile glands, non-glandular trichomes to 1 mm long; leaf indumentum not providing a hoary appearance"	18
18	"Stems with non-glandular trichomes to 1 mm and sparse, sessile glands 8-celled and dense; leaf with 6–11 pairs of teeth"	"Coleus apricus"
18	"Stems with non-glandular trichomes 0.2–0.5 mm and dense (15–35 trichomes/square mm), sessile glands 4-celled and sparse; leaf with (10–) 14–16 pairs of teeth"	"Coleus laxus"
19	"Stems and leaves with glandular trichomes very sparse or absent"	20
19	"Stems and/or leaves with glandular trichomes"	22
20	"Shrub to 80 cm; leaf adaxial surface hoary; verticillasters 10–20-flowered"	"Coleus gratus"
20	"Herb to 50 cm; leaf adaxial glossy; verticillasters 6–10-flowered"	21
21	"Stems with sessile glands sparse, non-glandular trichomes to 1 mm long; corolla tube angle <100º; fruiting calyx 4.5–5 mm long"	"Coleus laetus"
21	"Stems with sessile glands absent, non-glandular trichomes to 0.2 mm long; corolla tube angle >140º; fruiting calyx 3.6–3.8 mm long"	"Coleus nitidus"
22	"Stems, leaves and racemose inflorescences with dense glandular trichomes"	"Coleus dumicola"
22	"Stems, leaves and racemose inflorescences with scattered glandular trichomes"	23
23	"Non-glandular trichomes sparse to scattered, and up to 0.7 mm long, giving a glabrous appearance to the leaf; coloured sessile glands yellow"	24
23	"Non-glandular trichomes scattered to dense and up to 3 mm long, giving a felted appearance to the leaf; coloured sessile glands orange"	25
24	"Leaf with glandular trichomes and coloured sessile glands on abaxial surface; fruiting calyx 4.5–5.5 mm long"	"Coleus apreptus"
24	"Leaf without glandular trichomes and coloured sessile glands on abaxial surface; fruiting calyx 3–4.6 mm long"	"Coleus pulchellus"
25	"Leaves with 15–19 pairs of teeth; inflorescence axis with coloured sessile glands scattered; verticillasters 12–18-flowered; bracts broadly ovate 1.6–1.8 mm long"	"Coleus batianoffii"
25	"Leaves with 13–14 pairs of teeth; inflorescence axis with coloured sessile glands absent; verticillasters 8–14-flowered; bracts triangular-ovate 2.2–2.4 mm long"	"Coleus thalassoscopicus"
26	"Leaf lamina with 9–34 pairs of teeth"	27
26	"Leaf lamina with 4–10 pairs of teeth"	52
27	"Stems and lower leaf surfaces lacking glandular trichomes or with only sparse glandular trichomes in addition to the non-glandular trichomes"	28
27	"Stems and/or lower leaf surfaces with glandular trichomes scattered to densely covering in addition to the non-glandular trichomes"	40
28	"Stems and leaves both lacking or both very sparsely covered by coloured sessile glands"	29
28	"Stems and/or leaves (at least one) with coloured sessile glands dense"	32
29	"Leaves and branches succulent; strong savory aroma similar to oregano when touched or crushed; adaxial calyx lobe c. 1.5× length of lateral and abaxial lobes"	"Coleus amboinicus"
29	"Leaves and branches unpleasantly damp feeling (i.e. clammy) rather than succulent; aroma other than savory when touched or crushed; adaxial calyx lobe c. 1× length of lateral and abaxial lobes"	30
30	"Plant scented when crushed; inflorescence to 320 mm long, 1–11 branches; calyx indumentum antrorse"	"Coleus magnificus"
30	"Plant lacking scent when crushed; inflorescence <250 mm long, 1–4 branches; calyx indumentum divaricate to antrorse"	31
31	"Erect to decumbent shrub up to 100 cm tall with faint lemon smell when crushed; leaf lamina 10–80 × 10¬–65 mm, 10–19 pairs of teeth, trichomes < 2 mm long; flowers with orange-coloured sessile glands"	"Coleus bellus"
31	"Low growing herb to 30 cm tall with powerful sickly sweet aroma when touched; leaf lamina 10–40 mm × 6–25 mm, 7–14 pairs of teeth, trichomes < 0.8mm; flowers without orange-coloured sessile glands"	"Coleus fragrantissimus"
32	"Leaf lamina 2–2.5 times as long as wide; petiole 0.1–0.2 times as long as lamina"	"Coleus alloplectus"
32	"Leaf lamina 1–2 times as long as wide; petiole 0.2–0.5 times as long as lamina"	33
33	"Verticillasters 20–50-flowered; fruiting calyx 2–3 mm long"	"Coleus congestus"
33	"Verticillasters 1–22-flowered; fruiting calyx 3.5–5 mm long"	34
34	"Stems with only 4-celled coloured sessile glands on stems and leaves"	35
34	"Stems with coloured sessile glands on both stems and leaves but not 4-celled"	36
35	"Leaves with glandular trichomes absent; verticillasters 6–10-flowered; corolla tube slighly bent (0–30º) with coloured sessile glands and non-glandular trichomes absent, abaxial lobe 4–4.2 mm long"	"Coleus argentatus"
35	"Leaves with a few glandular trichomes present; verticillasters 14–16-flowered; corolla tube sharply bent (110–130º) with coloured sessile glands and non-glandular trichomes present, abaxial lobe 4.4–4.5 mm long"	"Coleus omissus"
36	"Pedicels 5–6 mm long; fruiting calyx >5.4 mm long"	"Coleus eungellaensis"
36	"Pedicels up to 4 mm long; fruiting calyx <5 mm long"	37
37	"Stems with coloured sessile glands scattered to dense, indumentum consisting of only 3–5-celled non-glandular trichomes 0.15–0.3 mm long; leaf adaxial surface with a few coloured sessile glands"	"Coleus amicorum"
37	"Stems lacking or with coloured sessile glands very sparse, indumentum with 6–10-celled non-glandular trichomes to 1.8 mm long present and 3–5-celled non-glandular trichomes ± present; leaf adaxial surface lacking coloured sessile glands"	38
38	"Pedicels 2–3 mm long; verticillasters 8–12-flowered; calyx lobes ‘flared’ (i.e. not held appressed to corolla tube); corolla pale lilac, anthers with white pollen"	"Coleus wallamanensis"
38	"Pedicels usually 0.1–2 mm long (rarely to 3.5 mm); verticillasters 12–22-flowered; calyx lobes held appressed to corolla tube; corolla blue, anthers with orange pollen"	39
39	"Stems with sessile glands ±present and glandular trichomes absent, non-glandular trichomes to 1.2 mm long; fruiting calyx with abaxial lobes 1.2–1.6 mm long"	"Coleus venustus"
39	"Stems with sessile glands and glandular trichomes very sparse, non-glandular trichomes to 1.8 mm long; fruiting calyx with abaxial lobes 2.2–2.4 mm long"	"Coleus magnificus"
40	"Coloured sessile glands very sparse to absent on branches"	41
40	"Coloured sessile glands scattered to dense on branches"	49
41	"Stems densely covered with glandular trichomes"	42
41	"Stems sparsely covered with or lacking glandular trichomes"	47
42	"Leaves stiffly succulent; strong savory aroma similar to oregano when touched or crushed; adaxial calyx lobe c. 1.5× length of lateral and abaxial lobes"	"Coleus amboinicus"
42	"Leaves not succulent or semi-succulent; aroma other than savory when touched or crushed; adaxial calyx lobe c. 1× length of lateral and abaxial lobes"	43
43	"Fruiting calyx (4)5–6 mm long, lateral and abaxial lobes longer than tube; corolla lacking coloured sessile glands"	"Coleus mirus"
43	"Fruiting calyx 2.5–5 mm long, lateral and abaxial lobes shorter than tube; corolla with coloured sessile glands"	44
44	"Inflorescence with coma and generally longer pedicels (to 4.5 mm); fruiting calyx < 3.3 mm long"	45
44	"Inflorescence without coma and generally shorter pedicels (1.7–2 mm); fruiting calyx > 3.3 mm long"	46
45	"Erect shrub to 300 cm tall covered in short non-glandular trichomes (<0.6 mm); leaves with 18–30 pairs of teeth; longer flowering calyx (2–2.5 mm) and fruiting calyx (3–3.3 mm long)"	"Coleus excelsus"
45	"Straggling to erect shrub to 100 cm tall covered in shaggy long non-glandular trichomes (up to 4 mm long); leaves with 10–14 pairs of teeth; shorter flowering calyx (1.5–1.6 mm) and fruiting calyx (2.5–2.6 mm)"	"Coleus geminatus"
46	"Leaves and Stems sometimes (usually at least at nodes) with some longer ± antrorse hairs; flowering calyx 2.3–3.3 mm long; fruiting calyx with adaxial lobe longer than lateral lobes and nearly as wide as to much narrower than long"	"Coleus diversus"
46	"Leaves and Stems sometimes with divaricate to retrorse hairs; flowering calyx 1.6–2.3 mm long; fruiting calyx with adaxial lobe not longer than lateral lobes and as wide as to much wider than long"	"Coleus graveolens"
47	"Plant with silver hairs; corolla tube with a 150º downward bend; fruiting calyx 5.4–6 mm long"	"Coleus torrenticola"
47	"Plant without silver hairs; corolla tube 120–130º downward bend; fruiting calyx 2.2–5.5 mm long"	48
48	"Plant covered with long non-glandular trichomes (to 2.3 mm); leaves with coloured sessile glands; inflorescence forming a coma"	"Coleus foetidus"
48	"Plant covered with short non-glandular trichomes (to 0.8 mm); leaves without coloured sessile glands; inflorescence not forming a coma"	"Coleus leiperi"
49	"Inflorescence with up to 11 branches, verticillasters 20–50-flowered; fruiting calyx < 3mm long"	"Coleus congestus"
49	"Inflorescence with up to 3 branches, verticillasters to 16-flowered; fruiting calyx > 3mm long"	50
50	"Stems and leaf nonglandular trichomes to 0.5 mm long, coloured sessile glands 4-celled; corolla tube with scattered non-glandular trichomes; fruiting calyx lateral lobe shorter than tube"	"Coleus altanmouiensis"
50	"Stems and/or leaf nonglandular trichomes to c. 1.6 mm long (at least at nodes), coloured sessile glands 4–8-celled; corolla tube glabrous; fruiting calyx lateral lobes equal to or longer than tube"	51
51	"Leaves with 16 or fewer pairs of teeth, coloured sessile glands only on abaxial surface, abaxial surface with non-glandular trichomes more dense on veins than lamina"	"Coleus amiculatus"
51	"Leaves with 16 or more pairs of teeth, coloured sessile glands present on both surfaces, abaxial surface with non-glandular trichomes less dense on veins than lamina"	"Coleus amoenus"
52	"Plant powerfully and unpleasantly aromatic when crushed, distinctively succulent; inflorescence bracts with a thickened, recurved apex; fruiting calyx densely covered by non-glandular trichomes in throat"	53
52	"Plant without a powerful unpleasant odour when crushed, not distinctively succulent; inflorescence bracts without a thickened recurved apex; fruiting calyx without non-glandular trichomes in throat"	54
53	"Flowers and fruits densely clustered throughout infloresence; corolla 8–10 mm long"	"Coleus caninus"
53	"Inflorescence usually with at least 1 or 2 spaced fruiting whorls below the flowers; corolla at least 12 mm long"	"Coleus neochilus"
54	"Coloured sessile glands on stems scattered to dense"	55
54	"Coloured sessile glands on stems absent to very few"	61
55	"Glandular trichomes absent on stems"	56
55	"Glandular trichomes present on stems"	59
56	"Leaves with thick silver grey indumentum on abaxial and adaxial surfaces; inflorescence with coma present in early development, verticillasters 12–14-flowered; flowers sessile or pedicels to 0.6 mm"	"Coleus fasciculatus"
56	"Leaves without silver-grey indumentum (or on one surface only); inflorescence without coma, main axis with coloured sessile glands, verticillasters with 10 or less flowers; pedicels 1–6 mm"	57
57	"Stems rounded, not tuberous, with 8–10-celled non-glandular trichomes up to 2 mm; leaves sweetly-scented and clammy"	"Coleus actites"
57	"Stems square, tuberous, with 1–4-celled non-glandular trichomes up to 0.2 mm; leaves faintly scented and not clammy"	58
58	"Stems densely covered with coloured sessile glands and without glandular trichomes; leaf laminae narrow ovate to narrow obovate; verticillasters 10–12-flowered; fruiting calyx > 5.7 mm long"	"Coleus insularis"
58	"Stems ± covered with coloured sessile glands and glandular trichomes; leaf laminae ovate, oblong ovate or sometimes orbicular; verticillasters 6–10-flowered; fruiting calyx < 5.6 mm long"	"Coleus australis"
59	"Stems with non-glandular trichomes up to 3 mm long, lacking glandular trichomes; calyx 1–1.6 mm long; fruiting calyx 2–3 mm long"	"Coleus congestus"
59	"Stems with non-glandular trichomes to 1.8 mm long, with glandular trichomes sparse to scattered; calyx 1.4–2.7 mm long; fruiting calyx 3.5–5.5 mm long"	60
60	"Erect subshrub to 100 cm high; stem with coloured sessile glands moderately dense, leaf densely covered by short hairs on abaxial and adaxial surface"	"Coleus intraterraneus"
60	"Erect to decumbent herb to 70 cm high; to semi erect herb prostrate lax semierect; stem with coloured sessile glands few or none; leaf moderately to sparsely covered by short hairs on abaxial and adaxial surface"	"Coleus australis"
61	"Non-glandular trichomes on branches dense"	62
61	"Non-glandular trichomes on branches sparse to absent"	67
62	"Leaf and calyx both without coloured sessile glands"	63
62	"Leaf and/or calyx (at least one) with coloured sessile glands"	64
63	"Foliage ± sweetly-scented when touched or crushed; leaves green with non-glandular trichomes long and shaggy (> 0.9 mm long); corolla < 9 mm long"	"Coleus habrophyllus"
63	"Strongly sweetly-scented when touched or crushed; leaves appearing silver with non-glandular trichomes closely appressed (< 0.6 mm long); corolla 9–12 mm long"	"Coleus fragrantissimus"
64	"Sweetly-scented when touched or crushed; leaves lacking coloured sessile glands"	"Coleus suaveolens"
64	"Lacking a sweet-scent when touched or crushed; leaves with coloured sessile glands present on abaxial surface"	65
65	"Shrub to 150 cm high; leaves to 110 mm long and 70 mm wide; with up to 11 racemose branches, axis densely covered by woolly long antrorse non-glandular trichomes"	"Coleus magnificus"
65	"Subshrub or herb to 70 cm high; leaves to 45 mm long and 42 mm wide; inflorescences with up to 5 racemose branches, axis covered by short to long (but not woolly) spreading non-glandular trichomes"	66
66	"Plant lacking scent when touched or crushed; stem thickened but without rounded tuberous base; inflorescence axis without coloured sessile glands, verticillasters 16–24-flowered; flowering calyx 2.6–2.8 mm long"	"Coleus acariformis"
66	"Plant with ± scent when touched or crushed (particularly main inflorescence axis); stem thickened into a fleshy tuberous base; inflorescence axis with coloured sessile glands; verticillasters (6–)10-flowered; flowering calyx 1.4–2.6 mm long"	"Coleus australis"
67	"Erect herb or subshrub to 100 cm; stem and leaves very densely covered by glandular trichomes"	68
67	"Erect to decumbent fleshy herb to 70 cm; stem and leaves with glandular trichomes absent to scattered"	69
68	"Stems with non-glandular trichomes sparse; leaves soft (not scabrid), lamina broad-ovate, 8–10 pairs of teeth; corolla lilac and 7–8 mm long"	"Coleus bipartitus"
68	"Stems with non-glandular trichomes generally absent; leaves scabrid, lamina broad-ovate, subcordate and/or reniform, 5–8 pairs of teeth; corolla deep purple and 10–11 mm long"	"Coleus splendens"
69	"Plant to 40 cm tall; seedling-derived stem not forming a fleshy tuberous base; living leaves dark green and appearing glabrous on adaxial surface, glandular trichomes absent; corolla glabrous"	"Coleus glabriflorus"
69	"Plant to 70 cm tall; seedling-derived stem forming a fleshy tuberous base; living leaves light green with non-glandular trichomes on adaxial surface visible, glandular trichomes ± present; non-glandular trichomes and coloured sessile glands present on corolla and calyx"	"Coleus australis"