1 "Fruiting heads grouped into conceptacles or clusters surrounded by ±fused bracts, often forming burrs, or spiny, sessile in the axils of leaves along the stems (male capitula sometimes in terminal or axillary ±spikelike clusters)" 2 1 "Fruiting heads not axillary conceptacles or burrs, or if with stiff barbed bristles, then in terminal heads" 6 2 "Leaves opposite, sometimes alternate above" 3 2 "Leaves alternate or in basal rosette" 5 3 "Leaves deeply divided" Ambrosia 3 "Leaves entire, toothed or rarely shallowly lobed" 4 4 "Fruits achenes with 2 terminal spines in heads surrounded by loose bracts" "Calyptocarpus {Calyptocarpus vialis}" 4 "Fruits achenes enclosed in spiny conceptacle of fused bracts" Acanthospermum 5 "Leaves deeply divided" Ambrosia 5 "Leaves entire, toothed, or if lobed or pinnately divided then fruits covered with hooked spines" Xanthium 6 "Flower heads with all florets with rays" 7 6 "Flower heads with a mixture of ray and tubular/filiform florets, or all tubular/filiform florets" 38 7 "Florets blue or violet, rarely white" 8 7 "Florets (at least outer ones) yellow to orange" 10 8 "Pappus of minute scales" "Cichorium {Cichorium intybus}" 8 "Pappus of plumose hairs (marginal achenes sometimes not plumose)" 9 9 "Leaves linear, grass-like, entire" Tragopogon 9 "Leaves oblong, margins toothed, sinuate or pinnatifid" Hypochaeris 10 "Receptacle paleate" 11 10 "Receptacle epaleate" 12 11 "Stems winged; plants spiny" Scolymus 11 "Stems without wings; soft herbs" Hypochaeris 12 "Involucral bracts in 1 whorl (2 whorls in Leontodon, but outer whorl tiny)" 13 12 "Involucral bracts in more than 1 whorl, usually gradate" 15 13 "Pappus of scales or bristles" Leontodon 13 "Pappus hairs plumose" 14 14 "Leaves oblanceolate to lyrate, margins toothed to pinnatisect" Urospermum 14 "Leaves linear, grass-like, entire" Tragopogon 15 "All leaves in a basal rosette" 16 15 "Leaves basal and/or cauline" 21 16 "Outer involucral bracts spreading or recurved at maturity" 17 16 "All bracts clasping and tightly appressed" 18 17 "Inner involucral bracts in several series; achene usually beaked" Taraxacum 17 "Inner involucral bracts in 2 series; achenes unbeaked" "Youngia {Youngia japonica}" 18 "Pappus of plumose hairs on central achenes only (scales on outer ones)" Leontodon 18 "Pappus of bristles or scales" 19 19 "Pappus of long scales or of bristles broader at the base" "Microseris {Microseris lanceolata}" 19 "Pappus of scabrid bristles which are not broader at base" 20 20 "Taprooted perennial herbs; achenes usually beaked" Taraxacum 20 "Stoloniferous herb, rooting at nodes; achenes not beaked" "Launaea {Launaea sarmentosa}" 21 "Achenes beaked apically" 22 21 "Achenes not beaked, abruptly constricted apically or gradually tapering" 27 22 "Pappus hairs plumose" 23 22 "Pappus hairs simple or at most minutely scabridulous" 24 23 "Outer involucral bracts conspicuously cordate to broadly ovate" "Helminthotheca {Helminthotheca echioides}" 23 "Outer involucral bracts not broadly ovate or cordate" Picris 24 "Achenes strongly compressed" Lactuca 24 "Achenes not compressed" 25 25 "Achene with 1 or more rings of teeth (scales) below beak" "Chondrilla {Chondrilla juncea}" 25 "Achenes without teeth below beak" 26 26 "Inner involucral bracts hardened & erect at maturity" Crepis 26 "All involucral bracts soft & reflexed at maturity" "Youngia {Youngia japonica}" 27 "Achenes strongly compressed" 28 27 "Achenes not or hardly compressed" 29 28 "Achenes elongate-elliptic" Sonchus 28 "Achenes flask-shaped" "Actites {Actites megalocarpus}" 29 "Pappus of plumose hairs" "Scorzonera {Scorzonera laciniata}" 29 "Pappus of bristles, scales or absent" 30 30 "Pappus absent" "Lapsana {Lapsana communis subsp. communis}" 30 "Pappus present, rarely absent and then achene 4-ribbed" 31 31 "Pappus of scales (inner achene pappus scales sometimes awned)" Leontodon 31 "Pappus of bristles or absent" 32 32 "Achenes with 6 to several ribs" 33 32 "Achenes rough and 4-ribbed or 4-angled" 37 33 "Stellate hairs present" 34 33 "Stellate hairs absent" 35 34 "Stolons absent; basal leaves with an abrupt transition from petiole to blade; upper stem and peduncles lacking long non-glandular hairs; achenes > 2.5 mm long, the ribs apically confluent in an obscure ring; pappus-bristles in two rows, the hairs unequal" "Hieracium {Hieracium murorum}" 34 "Leafy stolons present; basal leaves tapering gradually to base; achenes < 2.5 mm long, the ribs projecting distally to form a crenulate apex; pappus hairs in 1 row with a few shorter than the rest" Pilosella 35 "Stem leaves with clasping auricles" Crepis 35 "Stem leaves lacking clasping auricles" 36 36 "Pappus of numerous long bristles" "Youngia {Youngia japonica}" 36 "Pappus of a single row of minute bristles, inner achenes with additional 2–5 longer ones" Tolpis 37 "Pappus scale-like, broadened at base" Tolpis 37 "Pappus of fine silky hairs" Reichardia 38 "All florets tubular and/or filiform, or ray florets with short inconspicuous rays (<3 mm long); involucral bracts sometimes coloured and spreading, resembling rays" 39 38 "Outer florets with obvious rays (> 3mm long)" 250 39 "Styles with a tuft of hairs below the stigmatic branches; receptacles paleate with scales or bristles (receptacle with a single floret in Echinops); leaves usually spine-tipped (spiny thistles)" 40 39 "Styles lacking tuft of hairs below stigmatic branches; receptacles epaleate, or if with scales then leaves opposite and not spiny (Parthenium has alternate leaves, deciduous paleae; Pilbara has paleae and alternate leaves)" 59 40 "Receptacle epaleate, glabrous" 41 40 "Receptacle paleate with scales or setae" 42 41 "Heads simple, each an individual multi-floretted capitulum" Onopordum 41 "Heads compound, of many 1-floretted capitula" "Echinops {Echinops sphaerocephalus}" 42 "Receptacle with scales" 43 42 "Receptacle with setae" 48 43 "Floret corollas yellow" "Berkheya {Berkheya rigida}" 43 "Floret corollas purple, pink or white" 44 44 "Achenes cylindrical or 4-ribbed" 45 44 "Achenes compressed or flat" 46 45 "Achenes cylindrical; plants with latex" "Crupina {Crupina vulgaris}" 45 "Achenes 4-ribbed; plants lacking latex" "Hemisteptia {Hemisteptia lyrata}" 46 "Leaves scarcely lobed, lacking spines" Arctium 46 "Leaves lobed and spiny" 47 47 "Involucral bracts with a terminal spine only" Cirsium 47 "Involucral bracts with a terminal spine and 2–8 lateral spinules" "Picnomon {Picnomon acarna}" 48 "Involucral bracts leaf-like and loose" 49 48 "Involucral bracts scale-like and more or less appressed" 50 49 "Achenes compressed, flat" Centaurea 49 "Achenes 4-angled" Carthamus 50 "Achenes obliquely attached to receptacle" 51 50 "Achenes vertically attached to receptacle" 55 51 "Pappus deciduous" Rhapontium 51 "Pappus persistent, rarely absent" 52 52 "Pappus absent" Centaurea 52 "Pappus persistent" 53 53 "Achenes smooth or sulcate, never foveolate" Centaurea 53 "Achenes diversely ridged, ribbed or foveolate" 54 54 "Annual to 80 cm tall" Volutaria 54 "Perennial to 150 cm tall" "Mantisalca {Mantisalca salmantica}" 55 "Achenes compressed" 56 55 "Achenes terete;" 58 56 "Pappus 1–3.5 mm long" Arctium 56 "Pappus 15–40 mm long" 57 57 "Leaves uniformly green on upper surface" "Cynara {Cynara cardunculus subsp. flavescens}" 57 "Leaves white-veined or variegated on upper surface" "Silybum {Silybum marianum}" 58 "Pappus hairs simple or barbellate" Carduus 58 "Pappus hairs plumose" "Cynara {Cynara cardunculus subsp. flavescens}" 59 "Leaves opposite, at least towards base of plant" 60 59 "Leaves alternate (spirally arranged) or in a basal rosette" 113 60 "Leaves opposite at least basally, rarely in whorls" 61 60 "Leaves alternate and/or rosulate" 104 61 "Heads compound" 62 61 "Heads simple" 77 62 "Inflorescence loosely spike-like, with heads of 1–6 capitula each with 4 florets" "Pseudelephantopus {Pseudelephantopus spicatus}" 62 "Inflorescence a dense compound head" 63 63 "Corolla 4 lobed & split on one side" Elephantopus 63 "Corolla 5-lobed ±equisized (if fewer lobes then not split)" 64 64 "Secondary involucre (sometimes small) of densely clustered leaves and/or bracts enclosing compound heads" 65 64 "Secondary involucre absent, with just a few loosely arranged leaves below the compound head" 73 65 "Leaves opposite petiolate" "Asteraceae tribe Gnaphalieae" 65 "Leaves alternate radicle or opposite sessile" 66 66 "Compound head surrounded by numerous white or yellow scales" "Asteraceae tribe Gnaphalieae" 66 "Compound head surrounded by few scales or leaves" 67 67 "Receptacle paleate" "Asteraceae tribe Gnaphalieae" 67 "Receptacle epaleate" 68 68 "Corolla hairy" "Asteraceae tribe Gnaphalieae" 68 "Corolla glabrous" 69 69 "Outer florets female and/or all florets filiform" 70 69 "All florets tubular and bisexual" 71 70 "Plant stemless" "Asteraceae tribe Astereae" 70 "Plants with erect stems" "Asteraceae tribe Gnaphalieae" 71 "Pappus plumose, as long as or longer than corolla; plants prostrate" "Asteraceae tribe Gnaphalieae" 71 "Pappus absent, or of various types, but if as long as corolla, plants erect" 72 72 "Glabrous erect herb dichotomously branching from below terminal capitulum" Flaveria 72 "Usually hairy herbs, iregularly branching, lateral ininflorescences rarely overtopping terminal ones" "Asteraceae tribe Gnaphalieae" 73 "Individual capitulum with all florets alike (homogamous)" "Asteraceae tribe Gnaphalieae" 73 "Individual capitula with 2 types of florets (heterogamous)" 74 74 "Achenes tapering into a terminal beak" "Asteraceae tribe Gnaphalieae" 74 "Achenes more or less abruptly constricted below pappus" 75 75 "Stems not winged (if leaves decurrent, not along whole internode)" "Asteraceae tribe Gnaphalieae" 75 "Stems winged" 76 76 "Pappus bristle-like" Pterocaulon 76 "Pappus absent" Sphaeranthus 77 "Leaves lobed or deeply divided" 78 77 "Leaves entire or toothed" 82 78 "Pappus absent or of very short hairs or scales" 79 78 "Pappus hair-like, or with bristles longer than half the length of the achene" 80 79 "Water plant with soft branches; involucral bracts 4" Enydra 79 "Delicate terrestrial plant with stiffly erect branches; involucral bracts many" "Schkuhria {Schkuhria pinnata}" 80 "Upper leaves clasping stem; pappus of fine hairs" Emilia 80 "All leaves not clasping; pappus of few bristles" 81 81 "Involucral bracts fused into tube; plant with pungent smell" Tagetes 81 "Involucral bracts free; plant usually odourless" Bidens 82 "Pappus absent or very small" 83 82 "Pappus at least half as long as achene" 90 83 "Capitulum covered with glandular hairs" Sigesbeckia 83 "Capitulum without glands" 84 84 "Capitula axillary and sessile in axils of leaves" 85 84 "Capitula terminal and more or less stalked" 87 85 "Swamp plants; involucral bracts 4, outer 2 longer than inner two" Enydra 85 "Plants of dry land; involucral bracts 2 or many (never 4)" 86 86 "Florets 4 per capitulum; pappus of 2 awns fused to wings; involucral bracts 2" "Synedrellopsis {Synedrellopsis grisebachii}" 86 "Florets 7–50 per capitulum; pappus absent; involucral bracts many" "Iva {Iva axillaris}" 87 "Pappus absent" "Gymnocoronis {Gymnocoronis spilanthoides}" 87 "Pappus of small scales or a lacerate crown" 88 88 "Florets few (c. 5 or 6) per capitulum" "Eleutheranthera {Eleutheranthera ruderalis}" 88 "Florets numerous in each capitulum" 89 89 "Ray florets absent; disc florets usually blue to mauve, sometimes white" Ageratum 89 "Ray florets present, with white ligules to c. 1.8 mm long; disc florets whitish" Eclipta 90 "Capitula with 4 or 5 florets" 91 90 "Capitula with numerous florets" 92 91 "Pappus of numerous bristles" "Mikania {Mikania micrantha}" 91 "Pappus of 3 bristles alternating with 3 scales" Stevia 92 "Involucral bracts membranous" "Asteraceae tribe Gnaphalieae" 92 "Involucral bracts herbaceous, sometimes with membranous margins" 93 93 "Pappus hairs gland-tipped" Adenostemma 93 "Pappus hairs without glands" 94 94 "Leaves linear to oblong, entire, sometimes narrowly pinnatifid" 95 94 "Leaves (or leaflets) lanceolate to ovate, toothed or serrate, or entire" 99 95 "Involucral bracts broad" Acmella 95 "Involucral bracts linear to linear-lanceolate or oblong" 96 96 "Achenes flattened and winged" "Diodontium {Diodontium filifolium}" 96 "Achenes ±cylindrical, unwinged" 97 97 "Achenes with a long elongate beak, curled into an inverted J-shape" Glossocardia 97 "Achenes more or less truncate, not beaked" 98 98 "Awns spreading at c. 90º to body of achene, or reflexed" Trioncinia 98 "Awns erect or only slightly diverging" Glossocardia 99 "Leaves (at least upper ones) compound, trifoliate, entire" "Florestina {Florestina tripteris}" 99 "Leaves simple, toothed or serrate" 100 100 "Pappus of scales" Ageratum 100 "Pappus of hairs or bristles" 101 101 "Pappus of 2–4 bristles" "Calyptocarpus {Calyptocarpus vialis}" 101 "Pappus of many fine hairs" 102 102 "Perennial herbs; involucral bracts in 1 or 2 whorls" Eupatorium 102 "Shrubs; involucral bracts in 4–7 whorls" 103 103 "Leaves hirsute and gland-dotted on both surfaces; achenes attached laterally to conical receptacle" "Praxelis {Praxelis clematidea}" 103 "Leaves sparsely hairy above, more densely below, not gland-dotted; achenes attached terminally to flat receptacle" Chromolaena 104 "Pappus purplish-mauve" Erechtites 104 "Pappus white or almost so" 105 105 "Florets pink, or yellow-orange to orange, or reddish brown" 106 105 "Florets usually yellow or white (rarely pink or purple)" 108 106 "Leaves clasping upper stems; florets pink" Emilia 106 "Leaves not clasping stems; florets yellow-orange to reddish brown (sometimes drying pink)" 107 107 "Leaves more or less sessile, lacking a pair of stipule-like lobes" "Gynura {Gynura drymophila}" 107 "Leaves petiolate, lamina often with a pair of stipule-like lobes" "Crassocephalum {Crassocephalum crepidioides}" 108 "Annual herbs" Senecio 108 "Climbers, perennial herbs, shrubs or small trees" 109 109 "Climbers to 3 m" "Delairea {Delairea odorata}" 109 "Perennial herbs, shrubs or small trees" 110 110 "Lower leaf surface, peduncles & capitula densely woolly; shrubs or trees" Bedfordia 110 "Lower leaf surfaces, peduncles & capitula not densely woolly; herbs, or if shrubs or small trees, leaves glabrous" 111 111 "Leaves glabrous, viscid, gland-dotted; shrubs or small trees" Brachyglottis 111 "Leaves hairy, or if glabrous, not viscid and gland-dotted; perennial herbs" 112 112 "Capitula disciform; outer florets with a rudimentary ligule c. 1 mm long; central bisexual or functionally male florets 2–5" "Arrhenechthites {Arrhenechthites mixtus}" 112 "Capitula discoid, or if disciform then outer florets without a ligule; central bisexual florets usually more than 5" Senecio 113 "Involucral bracts ±equisized and fused in a single whorl, sometimes with a few very short bracts at base; flower heads simple capitula" 114 113 "Involucral bracts usually gradated, in 2 or more whorls and free, or flower heads compound" 167 114 "Leaves opposite at least basally, rarely in whorls" 115 114 "Leaves alternate and/or rosulate" 158 115 "Heads compound" 116 115 "Heads simple" 131 116 "Inflorescence loosely spike-like, with heads of 1–6 capitula each with 4 florets" "Pseudelephantopus {Pseudelephantopus spicatus}" 116 "Inflorescence a dense compound head" 117 117 "Corolla 4 lobed & split on one side" Elephantopus 117 "Corolla 5-lobed ±equisized (if fewer lobes then not split)" 118 118 "Secondary involucre (sometimes small) of densely clustered leaves and/or bracts enclosing compound heads" 119 118 "Secondary involucre absent, with just a few loosely arranged leaves below the compound head" 127 119 "Leaves opposite and petiolate" "Asteraceae tribe Gnaphalieae" 119 "Leaves alternate and radicle, or opposite and sessile" 120 120 "Compound head surrounded by numerous white or yellow scales" "Asteraceae tribe Gnaphalieae" 120 "Compound head surrounded by few scales or leaves" 121 121 "Receptacle paleate" "Asteraceae tribe Gnaphalieae" 121 "Receptacle epaleate" 122 122 "Corolla hairy" "Asteraceae tribe Gnaphalieae" 122 "Corolla glabrous" 123 123 "Outer florets female and/or all florets filiform" 124 123 "All florets tubular and bisexual" 125 124 "Plant stemless" "Asteraceae tribe Astereae" 124 "Plants with erect stems" "Asteraceae tribe Gnaphalieae" 125 "Pappus plumose, as long as or longer than corolla; plants prostrate" "Asteraceae tribe Gnaphalieae" 125 "Pappus absent, or of various types, but if as long as corolla, plants erect" 126 126 "Glabrous erect herb dichotomously branching from below terminal capitulum" Flaveria 126 "Usually hairy herbs, irregularly branching, lateral ininflorescences rarely overtopping terminal ones" "Asteraceae tribe Gnaphalieae" 127 "Individual capitulum with all florets alike (homogamous)" "Asteraceae tribe Gnaphalieae" 127 "Individual capitula with 2 types of florets (heterogamous)" 128 128 "Achenes tapering into a terminal beak" "Asteraceae tribe Gnaphalieae" 128 "Achenes more or less abruptly constricted below pappus" 129 129 "Stems not winged (if leaves decurrent, not along whole internode)" "Asteraceae tribe Gnaphalieae" 129 "Stems winged" 130 130 "Pappus bristle-like" Pterocaulon 130 "Pappus absent" Sphaeranthus 131 "Leaves lobed or deeply divided" 132 131 "Leaves entire or toothed" 136 132 "Pappus absent or of very short hairs or scales" 133 132 "Pappus hair-like, or with bristles longer than half the length of the achene" 134 133 "Water plant with soft branches; involucral bracts 4" Enydra 133 "Delicate terrestrial plant with stiffly erect branches; involucral bracts many" "Schkuhria {Schkuhria pinnata}" 134 "Upper leaves clasping stem; pappus of fine hairs" Emilia 134 "All leaves not clasping; pappus of few bristles" 135 135 "Involucral bracts fused into tube; plant with pungent smell" Tagetes 135 "Involucral bracts free; plant usually odourless" Bidens 136 "Pappus absent or very small" 137 136 "Pappus at least half as long as achene" 144 137 "Capitulum covered with glandular hairs" Sigesbeckia 137 "Capitulum without glands" 138 138 "Capitula axillary and sessile in axils of leaves" 139 138 "Capitula terminal and more or less stalked" 141 139 "Swamp plants; involucral bracts 4, outer 2 longer than inner two" Enydra 139 "Plants of dry land; involucral bracts 2 or many (never 4)" 140 140 "Florets 4 per capitulum; pappus of 2 awns fused to wings; involucral bracts 2" "Synedrellopsis {Synedrellopsis grisebachii}" 140 "Florets 7–50 per capitulum; pappus absent; involucral bracts many" "Iva {Iva axillaris}" 141 "Pappus absent" "Gymnocoronis {Gymnocoronis spilanthoides}" 141 "Pappus of small scales or a lacerate crown" 142 142 "Florets few (c. 5 or 6) per capitulum" "Eleutheranthera {Eleutheranthera ruderalis}" 142 "Florets numerous in each capitulum" 143 143 "Ray florets absent; disc florets usually blue to mauve, sometimes white" Ageratum 143 "Ray florets present, with white ligules to c. 1.8 mm long; disc florets whitish" Eclipta 144 "Capitula with 4 or 5 florets" 145 144 "Capitula with numerous florets" 146 145 "Pappus of numerous bristles" "Mikania {Mikania micrantha}" 145 "Pappus of 3 bristles alternating with 3 scales" Stevia 146 "Involucral bracts membranous" "Asteraceae tribe Gnaphalieae" 146 "Involucral bracts herbaceous, sometimes with membranous margins" 147 147 "Pappus hairs gland-tipped" Adenostemma 147 "Pappus hairs without glands" 148 148 "Leaves linear to oblong, entire, sometimes narrowly pinnatifid" 149 148 "Leaves (or leaflets) lanceolate to ovate, toothed or serrate, or entire" 153 149 "Involucral bracts broad" Acmella 149 "Involucral bracts linear to linear-lanceolate or oblong" 150 150 "Achenes flattened and winged" "Diodontium {Diodontium filifolium}" 150 "Achenes ±cylindrical, unwinged" 151 151 "Achenes with a long elongate beak, curled into an inverted J-shape" Glossocardia 151 "Achenes more or less truncate, not beaked" 152 152 "Awns spreading at c. 90º to body of achene, or reflexed" Trioncinia 152 "Awns erect or only slightly diverging" Glossocardia 153 "Leaves (at least upper ones) compound, trifoliate, entire" "Florestina {Florestina tripteris}" 153 "Leaves simple, toothed or serrate" 154 154 "Pappus of scales" Ageratum 154 "Pappus of hairs or bristles" 155 155 "Pappus of 2–4 bristles" "Calyptocarpus {Calyptocarpus vialis}" 155 "Pappus of many fine hairs" 156 156 "Perennial herbs; involucral bracts in 1 or 2 whorls" Eupatorium 156 "Shrubs; involucral bracts in 4–7 whorls" 157 157 "Leaves hirsute and gland-dotted on both surfaces; achenes attached laterally to conical receptacle" "Praxelis {Praxelis clematidea}" 157 "Leaves sparsely hairy above, more densely below, not gland-dotted; achenes atached terminally to flat receptacle" Chromolaena 158 "Pappus purplish-mauve" Erechtites 158 "Pappus white or almost so" 159 159 "Florets pink, or yellow-orange to orange, or reddish brown" 160 159 "Florets usually yellow or white (rarely pink or purple)" 162 160 "Leaves clasping upper stems; florets pink" Emilia 160 "Leaves not clasping stems; florets yellow-orange to reddish brown (sometimes drying pink)" 161 161 "Leaves more or less sessile, lacking a pair of stiplule-like lobes" "Gynura {Gynura drymophila}" 161 "Leaves petiolate, lamina often with a pair of stipule-like lobes" "Crassocephalum {Crassocephalum crepidioides}" 162 "Annual herbs" Senecio 162 "Climbers, perennial herbs, shrubs or small trees" 163 163 "Climbers to 3 m" "Delairea {Delairea odorata}" 163 "Perennial herbs, shrubs or small trees" 164 164 "Lower leaf surface, peduncles & capitula densely woolly; shrubs or trees" Bedfordia 164 "Lower leaf surfaces, peduncles & capitula not densely woolly" 165 165 "Leaves glabrous, viscid, gland-dotted; shrubs or small trees" Brachyglottis 165 "Leaves hairy, or if glabrous, not viscid and gland-dotted; perennial herbs" 166 166 "Capitula disciform; outer florets with a rudimentary ligule c. 1 mm long; central bisexual or functionally male florets 2–5" "Arrhenechthites {Arrhenechthites mixtus}" 166 "Capitula discoid, or if disciform then outer florets without a ligule; central bisexual florets usually more than 5" Senecio 167 "Flower heads compound, with capitula tightly clustered and densely surrounded by scales and leaves, rarely scales and leaves absent" 168 167 "Flower heads not as above (usually of single capitula), sessile or stalked" 183 168 "Inflorescence loosely spike-like, with heads of 1–6 capitula each with 4 florets" "Pseudelephantopus {Pseudelephantopus spicatus}" 168 "Inflorescence a dense compound head" 169 169 "Corolla 4 lobed & split on one side" Elephantopus 169 "Corolla 5-lobed ±equisized (if fewer lobes then not split)" 170 170 "Secondary involucre (sometimes small) of densely clustered leaves and/or bracts enclosing compound heads" 171 170 "Secondary involucre absent, with just a few loosely arranged leaves below the compound head" 179 171 "Leaves opposite petiolate" "Asteraceae tribe Gnaphalieae" 171 "Leaves alternate radicle or opposite sessile" 172 172 "Compound head surrounded by numerous white or yellow scales" "Asteraceae tribe Gnaphalieae" 172 "Compound head surrounded by few scales or leaves" 173 173 "Receptacle paleate" "Asteraceae tribe Gnaphalieae" 173 "Receptacle epaleate" 174 174 "Corolla hairy" "Asteraceae tribe Gnaphalieae" 174 "Corolla glabrous" 175 175 "Outer florets female and/or all florets filiform" 176 175 "All florets tubular and bisexual" 177 176 "Plant stemless" "Asteraceae tribe Astereae" 176 "Plants with erect stems" "Asteraceae tribe Gnaphalieae" 177 "Pappus plumose, as long as or longer than corolla; plants prostrate" "Asteraceae tribe Gnaphalieae" 177 "Pappus absent, or of various types, but if as long as corolla, plants erect" 178 178 "Glabrous erect herb dichotomously branching from below terminal capitulum" Flaveria 178 "Usually hairy herbs, iregularly branching, lateral ininflorescences rarely overtopping terminal ones" "Asteraceae tribe Gnaphalieae" 179 "Individual capitulum with all florets alike (homogamous)" "Asteraceae tribe Gnaphalieae" 179 "Individual capitula with 2 types of florets (heterogamous)" 180 180 "Achenes tapering into a terminal beak" "Asteraceae tribe Gnaphalieae" 180 "Achenes more or less abruptly constricted below pappus" 181 181 "Stems not winged (if leaves decurrent, not along whole internode)" "Asteraceae tribe Gnaphalieae" 181 "Stems winged" 182 182 "Pappus bristle-like" Pterocaulon 182 "Pappus absent" Sphaeranthus 183 "Involucral bracts herbaceous, rarely with membranous margins, and tip appressed" 184 183 "At least some (often all) involucral bracts dry, papery and/or membranous or cartilagineous, and apices often recurved or spreading, mimicing ray florets" 221 184 "Pappus absent or cup-shaped (absent on achenes)" 185 184 "Pappus present" 206 185 "Achenes beaked" 186 185 "Achenes not beaked, more or less abruptly constricted towards apex" 187 186 "Achenes cylindrical; plants erect" "Asteraceae tribe Gnaphalieae" 186 "Achenes compressed; plants with basal rosette, rarely almost erect" "Asteraceae tribe Astereae" 187 "Ray floret corollas blue to purple" 188 187 "Ray floret corollas yellow or white, rarely tinged red, or lacking ray florets (Centipeda, Abrotanella)" 191 188 "Achenes smooth" "Asteraceae tribe Astereae" 188 "Achenes ribbed or angled" 189 189 "Achenes usually 4-angled or ribbed" "Ethulia {Ethulia conyzoides}" 189 "Achenes with 8–10 ribs" 190 190 "Outer involucral bracts leaf-like & longer than florets" Centratherum 190 "All involucral bracts shorter than florets" Pleurocarpaea 191 "Involucral bracts leaf-like; florets subtended by paleae" "Asteraceae tribe Gnaphalieae" 191 "Involucral bracts herbaceous; paleae minute or absent" 192 192 "Achenes terete or 3- or more angled" 193 192 "Achenes compressed but often ribbed" 200 193 "Leaves entire or almost so (minutely denticulate in Abrotonella forsteroides)" 194 193 "Leaves toothed or deeply divided" 196 194 "All florets bisexual" "Asteraceae tribe Gnaphalieae" 194 "Outer florets female; inner florets bisexual or male" 195 195 "Inner florets bisexual" Centipeda 195 "Inner florets male" Abrotanella 196 "Leaves with a few teeth towards apex or evenly serrate, but never deeply divided" Centipeda 196 "Leaves deeply divided" 197 197 "Plants usually >50 cm tall" Artemisia 197 "Plants usually <50 cm tall" 198 198 Subshrubs Pentzia 198 "Annual herbs" 199 199 "Plants glabrous; disc greenish" Matricaria 199 "Plants sparsely hairy; disc yellow" Oncosiphon 200 "Outer florets with short rays" "Eriocephalus {Eriocephalus africanus}" 200 "Outer florets tubular or filiform" 201 201 "All florets tubular" Abrotanella 201 "Outer florets filiform or reduced" 202 202 "Pappus of bristle-like hairs" Blumea 202 "Pappus absent or minute" 203 203 "Capitula sessile, achenes spinous apically" 204 203 "Capitula on distinct peduncle; achenes lacking spines" 205 204 "Achenes with 2 ciliate horns on both sides of apex" "Asteraceae tribe Astereae" 204 "Achenes with 2 spiny wings and spinescent syle" Soliva 205 "All leaves in basal rosette" "Asteraceae tribe Astereae" 205 "Leaves basal and cauline" Cotula 206 "Involucral bracts leaf-like and spreading" "Asteriscus {Asteriscus spinosus}" 206 "Involucral bracts short and more or less appressed" 207 207 "Pappus of at least fertile flowers scale-like" 208 207 "Pappus hair-like, sometimes barbed" 209 208 "Florets tubular, 5-lobed" "Asteraceae tribe Gnaphalieae" 208 "Outer florets filiform, 2- or 3-lobed" "Asteraceae tribe Astereae" 209 "Pappus rose or pink" "Asteraceae tribe Gnaphalieae" 209 "Pappus white or yellow but florets sometimes purple" 210 210 "Outer florets filiform (sometimes few)" 211 210 "Outer florets various but not filiform" 212 211 "Leaves toothed or lobed, rarely entire; filiform florets numerous" Blumea 211 "Leaves entire; filiform florets few" "Asteraceae tribe Gnaphalieae" 212 "Achenes beaked" "Asteraceae tribe Gnaphalieae" 212 "Achenes not beaked, abruptly constricted towards apex" 213 213 "All florets tubular" 214 213 "Outer florets filiform or with rays" 218 214 "Achenes 5–10 ribbed" 215 214 "Achenes lacking ribs" 217 215 "Leaves coarsely biserrate; stiffly erect shrubs" Pilbara 215 "Leaves entire or almost so; perennial herbs or scrambling shrubs" 216 216 "Perennial herbs; achenes inconspicuously 4-5-ribbed" Vernonia 216 "Scrambling shrubs; achenes 10-ribbed" Decaneuropsis 217 "Annual or perennial taprooted herbs or short-lived shrubs" Cyanthillium 217 "Rhizomatous shrubby climbers" "Tarlmounia {Tarlmounia elliptica}" 218 "Outer achenes flat, inner ones terete" "Asteraceae tribe Astereae" 218 "All achenes similar or aborted" 219 219 "Fruits flat" "Asteraceae tribe Astereae" 219 "Fruits terete, rarely slightly compressed" 220 220 "Involucral bracts linear; herbs" "Asteraceae tribe Gnaphalieae" 220 "Involucral bracts ovate with dry margins; shrubs or subshrubs" "Asteraceae tribe Astereae" 221 "Achenes with two thick spines apically; pappus absent" "Asteraceae tribe Gnaphalieae" 221 "Achenes lacking spines, or spines slender; pappus usually present" 222 222 "Pappus absent, cup-shaped or much reduced" 223 222 "Pappus of hairs, bristles or scales" 230 223 "Corolla and whole plant glandular" "Asteraceae tribe Gnaphalieae" 223 "Corolla glabrous or rarely leaves glandular" 224 224 "Leaves deeply divided" Tanacetum 224 "Leaves entire or toothed" 225 225 "Achenes beaked" "Carpesium {Carpesium cernuum}" 225 "Achenes more or less abruptly constricted apically" 226 226 "Involucral bracts broadly ovate, apices not spreading; paleae equalling or exceeding florets" "Asteraceae tribe Gnaphalieae" 226 "Involucral bracts usually lanceolate to elliptic, apices spreading or not; paleae absent or usually shorter than florets" 227 227 "Receptacle paleate" "Asteraceae tribe Gnaphalieae" 227 "Receptacle epaleate" 228 228 "All florets alike" "Asteraceae tribe Gnaphalieae" 228 "Outer florets filiform" 229 229 "Filiform florets few; leaves entire" "Asteraceae tribe Gnaphalieae" 229 "Filiform florets numerous; at least some leaves toothed or lobed" Epaltes 230 "Achenes abruptly constricted into neck or beak below pappus" 231 230 "Achenes lacking neck or beak below pappus" 233 231 "Involucral bracts with petaloid spreading tips" "Asteraceae tribe Gnaphalieae" 231 "Involucral bracts appressed and membranous" 232 232 "Plants with glandular hairs, aromatic; capitula sessile" Dittrichia 232 "Plants lacking glandular hairs; capitula pedunculate" "Asteraceae tribe Gnaphalieae" 233 "Middle involucral bracts pouched" "Asteraceae tribe Gnaphalieae" 233 "No bracts pouched" 234 234 "Plants with 2 types of capitula (tubular florets, tubular and/or filiform)" 235 234 "Plants with all capitula similar" 236 235 "Mat- or hummock-forming plants, capitula cup-shaped" "Asteraceae tribe Gnaphalieae" 235 "Shrubs or subshrubs; capitula cylindrical" Cratystylis 236 "Outer florets with short rays; achene shed in complex structure including paleae and bracts" "Parthenium {Parthenium hysterophorus}" 236 "Outer florets lacking rays; achenes shed independently of paleae and bracts" 237 237 "All florets in capitulum filiform" "Asteraceae tribe Gnaphalieae" 237 "All florets in capitulum either tubular or filiform & tubular" 238 238 "Outer slender or filiform florets rare or absent" 239 238 "Outer filiform florets in several rows" 241 239 "Pappus hair-like, simple, barbed or plumose" "Asteraceae tribe Gnaphalieae" 239 "Pappus scaly at least at base, if hair-like and plumose, then inner involucral bracts linear-lanceolate, pointed" 240 240 "Florets all similar and bisexual" "Asteraceae tribe Gnaphalieae" 240 "Outer florets filiform, female" "Coleocoma {Coleocoma centaurea}" 241 "Leaves obovate spathulate, 4 mm long" "Asteraceae tribe Gnaphalieae" 241 "Leaves linear-lanceolate, oblanceolate, usually more than 8 mm long" 242 242 "Perennials, usually shrubs or subshrubs" 243 242 "Annual herbs, rarely with slightly woody base" 245 243 "Pappus of narrow scales" "Thespidium {Thespidium basiflorum}" 243 "Pappus of hairs" 244 244 "Mat-forming alpine plants" "Asteraceae tribe Gnaphalieae" 244 "Erect plants of tropics & subtropics" Pluchea 245 "Pappus absent on filiform outer florets" Epaltes 245 "Pappus present on all florets" 246 246 "Pappus of narrow scales" "Coleocoma {Coleocoma centaurea}" 246 "Pappus of simple or plumose hairs" 247 247 "Pappus hairs simple" "Asteraceae tribe Gnaphalieae" 247 "Pappus hairs barbed or plumose on fertile florets" 248 248 "Florets few (c. 6), disc florets sterile" "Allopterigeron {Allopterigeron filifolius}" 248 "Florets numerous (15–190; disc florets fertile" 249 249 "Mat-forming alpine plants, leaves not noticeably aromatic" "Asteraceae tribe Gnaphalieae" 249 "Erect plants of tropics & subtropics; leaves often aromatic" Streptoglossa 250 "Leaves opposite, at least towards base of plant" 251 250 "Leaves alternate (spirally arranged) and/or in a basal rosette" 282 251 "Bracts and whole plant covered with glandular hairs" 252 251 "Bracts and whole plant without glandular hairs" 253 252 "Leaves linear, sessile, opposite below, alternate above" "Madia {Madia sativa}" 252 "Leaves lanceolate, elliptic, petiolate, opposite" Sigesbeckia 253 "Corolla of outer florets longer than (7–) 8 mm" 254 253 "Corolla of outer florets up to 5 mm long (4–7 mm in Centromadia {Centromadia pungens})" 268 254 "Leaves entire, toothed or shallowly lobed" 255 254 "Leaves (at least lower ones) deeply lobed" 265 255 "Leaves sessile and clasping stem" 256 255 "Leaves petiolate or constricted towards base, not clasping" 257 256 "Leaves usually entire; involucral bracts in at least 2 whorls" Zinnia 256 "Leaves usually toothed; involucral bracts in 2 whorls" "Guizotia {Guizotia abyssinica}" 257 "Pappus of many hairs" 258 257 "Pappus of few hairs, absent, or a cup" 259 258 "Plants prostrate or procumbent perennial herb" "Tridax {Tridax procumbens}" 258 "Plants erect shrubs or subshrubs" "Asteraceae tribe Astereae" 259 "Achenes winged" "Verbesina {Verbesina encelioides}" 259 "Achenes angled or compressed but without wings" 260 260 "Pericarp thickened and extended above pappus, forming a cup" "Sphagneticola {Sphagneticola trilobata}" 260 "Pericarp if thickened, sometimes forming wings, but not a cup, or not thickened" 261 261 "Paleae blunt, just exceeding achenes and ±hooded" Wollastonia 261 "Paleae acute, shorter than or exceeding achenes, never hooded" 262 262 "Paleae soft, membranous" 263 262 "Paleae stiff, cartilaginous" 264 263 "Receptacle conical" Acmella 263 "Receptacle flat to convex" "Pascalia {Pascalia glauca}" 264 "Outer involucral bracts longer than inner ones" Acunniana 264 "Outer involucral bracts shorter than or equalling inner ones" Apowollastonia 265 "Lobes of leaves up to 2 mm wide" Cosmos 265 "Lobes of leaves 4–50 mm wide" 266 266 "Plants subshrubs up to 80 cm tall; ray corollas yellow" Coreopsis 266 "Plants up to 4 m tall; floret corollas white, yellow, red or purplish" 267 267 "Paleae accrescent; corollas white" Montanoa 267 "Paleae not accrescent; corollas various colours incl. white" Dahlia 268 "Receptacle epaleate" 269 268 "Receptacle paleate" 270 269 "Involucral bracts in 1 whorl and fused; plant with pungent smell" Tagetes 269 "Involucral bracts in several whorls; free; not pungently aromatic" "Schkuhria {Schkuhria pinnata}" 270 "Pappus of awns, bristles or scales" 271 270 "Pappus absent, rarely a cup, or very short hairs or scales" 273 271 "Leaves pinnately divided (at least basally)" Bidens 271 "Leaves entire or toothed" 272 272 "Achenes of 2 types: ray achenes flattened with short marginal deeply lacerate wing; disc achenes biconvex with 2 apical awns" "Synedrella {Synedrella nodiflora}" 272 "Achenes all similar, obconic; pappus of numerous fimbriate scales" "Galinsoga {Galinsoga parviflora}" 273 "Involucral bracts covered with stalked glands" Sigesbeckia 273 "Involucral bracts without glands" 274 274 "Tubular florets sterile;" 275 274 "All florets fertile; involucral bracts numerous in 2 or more whorls" 277 275 "Involucral bracts many; achene with offset apical horn" "Centromadia {Centromadia pungens}" 275 "Involucral bracts 4 to 6; achenes lacking apical horn" 276 276 "Capitula sessile in leaf axils; ray florets in several whorls" Enydra 276 "Capitula stalked (sometimes shortly); ray florets (always 5) in 1 whorl" Pentalepis 277 "Receptacle conical" Acmella 277 "Receptacle flat or slightly convex" 278 278 "Paleae blunt or truncate" 279 278 "Paleae acute" 280 279 "Paleae truncate, erose, membranous" Blainvillea 279 "Paleae blunt, not erose, cartilaginous" Wollastonia 280 "Outer achenes 3-angled in section, inner ones 2-angled, often winged" Eclipta 280 "All achenes similar, sometimes angled, rarely winged" 281 281 "Outer involucral bracts longer that inner ones" Acunniana 281 "Outer involucral bracts shorter than or equalling inner ones" Apowollastonia 282 "Pappus present and usually prominent" 283 282 "Pappus absent, reduced to less than 1 mm, or a cup" 316 283 "Leaves linear in clusters along stem" 284 283 "Leaves alternate, cauline, if clustered then not linear" 285 284 "Pappus of 2 awns" "Diodontium {Diodontium filifolium}" 284 "Pappus of 5 translucent, usually awned, scales" "Helenium {Helenium amarum var. amarum}" 285 "Involucral bracts in 1 whorl, usually fused, rarely with a few much shorter ones basally" 286 285 "Involucral bracts in 2 to several whorls, free" 292 286 "Pappus scale-like" Gazania 286 "Pappus of hairs or bristles" 287 287 "Capitula borne singly on long peduncles in lateral positions" Euryops 287 "Capitula usually in terminal clusters, rarely solitary and terminal" 288 288 "Shrubs to 3.5 m; leaves viscid, adaxial (upper) surface gland-dotted" Brachyglottis 288 "Herbs or shrubs; leaves not viscid or gland-dotted" 289 289 "Leaves with distinct lamina & petiole; petiole 5 cm or more long" 290 289 "Leaves sessile, rarely petiolate, and then petiole much less than 5 cm long" 291 290 "Hermaphrodite shrubs to 3 m high; ligules yellow" "Roldana {Roldana petasitis}" 290 "Rosetted, dioecious herbs to 0.4 m high (plants male in Australia); ligules white" "Petasites {Petasites pyrenaicus}" 291 "Achenes compressed" "Cineraria {Cineraria lyratiformis}" 291 "Achenes not compressed" Senecio 292 "Receptacle paleate" 293 292 "Receptacle rough or pitted but lacking paleae" 300 293 "Involucral bracts leaf-like and much longer than florets" "Asteriscus {Asteriscus spinosus}" 293 "Involucral bracts shorter than or equalling florets" 294 294 "Plants usually exceeding 1 m tall, and with 1–several erect branches" 295 294 "Plants with basal rootstock, somewhat branched; branches up to 50 cm" 297 295 "Leaves deeply divided" Tithonia 295 "Leaves entire or toothed" 296 296 "Pappus of short bristles; rays lobed or distinctly toothed" "Verbesina {Verbesina encelioides}" 296 "Pappus of deciduous scales; rays scarcely toothed" Helianthus 297 "Pappus of 2 deciduous scales" Helianthus 297 "Pappus of 2 persistent awns, or of 6–10 persistent scales" 298 298 "Rays large and spectacular" Gaillardia 298 "Rays scarcely longer than involucre" 299 299 "Achenes flattened and winged" "Diodontium {Diodontium filifolium}" 299 "Achenes cylindrical" Glossocardia 300 "Involucral bracts fused into a tube basally" Gazania 300 "Involucral bracts in several whorls, free" 301 301 "At least some leaves divided or lobed" 302 301 "Leaves entire and linear, or toothed" 305 302 "Ray florets sterile" 303 302 "Ray florets female" 304 303 "Outer pappus obovate scale, inner pappus bristles" Ursinia 303 "All pappus small scales" Arctotheca 304 "Achenes with 3 wing-like ridges on 1 side; pappus of scales" "Arctotis {Arctotis stoechadifolia}" 304 "Achenes ribbed but not winged; pappus of bristles" "Asteraceae tribe Astereae" 305 "Inner involucral bracts dry and membranous" 306 305 "All involucral bracts herbaceous, or sometimes with membranous margins" 308 306 "Achenes with a short neck below pappus" Dittrichia 306 "Achenes cylindrical, without a neck" 307 307 "Style branches slender and pointed" "Allopterigeron {Allopterigeron filifolius}" 307 "Style branches short and blunt" "Asteraceae tribe Gnaphalieae" 308 "Fruits beaked" "Asteraceae tribe Astereae" 308 "Fruits not beaked" 309 309 "Ray florets in 2 or more whorls" 310 309 "Ray florets in 1 whorl" 311 310 "Involucral bracts and receptacle glandular; ligules 2-4-lobed" Streptoglossa 310 "Involucral bracts and receptacle not glandular; ligules entire" "Asteraceae tribe Astereae" 311 "Pappus a double row of brick red bristles, outer short, inner long" "Asteraceae tribe Astereae" 311 "Pappus 1-seriate or if a double row of bristles, not brick red and of 2 lengths" 312 312 "All florets fertile (inner disc florets sterile in Grindelia)" 313 312 "Either disc or ray florets sterile" 315 313 "Pappus of 2-10 simple caducous awns" "Asteraceae tribe Astereae" 313 "Pappus usually of barbellate or plumose bristles or awns or absent" 314 314 "Ray florets with staminodes" Trichocline 314 "Ray florets without staminodes" "Asteraceae tribe Astereae" 315 "Ray florets female; disc florets usually staminate; pappus of scales, or rarely plumose bristles or absent (pseudo-pappus of barbellate awns derived from pericarp)" "Asteraceae tribe Astereae" 315 "Ray florets neuter; disc florets bisexual; pappus and pseudo-pappus not as above" Arctotheca 316 "Plants tufted, with linear leaves, old leaves clothing base" "Asteraceae tribe Astereae" 316 "Plants erect, or if with a basal rosette, then leaves not linear" 317 317 "Achenes with a beak" 318 317 "Achenes abruptly constricted at apex, not beaked or rostrate" 319 318 "Achenes polymorphic, some rostrate; rays yellow" "Oligocarpus {Oligocarpus calendulaceus}" 318 "Achenes monomorphic, all shortly beaked; rays white to purple" "Asteraceae tribe Astereae" 319 "Involucral bracts with a terminal spine" 320 319 "Involucral bracts without terminal spine" 321 320 "Perennials; leaves with spines" "Berkheya {Berkheya rigida}" 320 "Annuals; leaves without spines" "Gorteria {Gorteria personata}" 321 "Receptacle paleate" 322 321 "Receptacle epaleate" 328 322 "Capitula small and in terminal clusters" Achillea 322 "Capitula larger and individually pedunculate or solitary" 323 323 "Leaves lobed to almost entire" Cymbonotus 323 "Leaves deeply divided" 324 324 "Paleae longer than outer florets" "Asteraceae tribe Astereae" 324 "Paleae shorter than florets" 325 325 "Achenes woolly" "Lasiospermum {Lasiospermum bipinnatum}" 325 "Achenes glabrous" 326 326 "Rhizomatous perennials; achenes 3-ribbed" "Chamaemelum {Chamaemelum nobile}" 326 "Annuals or perennials, not rhizomatous; achenes c. 8–10-ribbed" 327 327 "Ligules yellow; pappus a small corona" "Cota {Cota tinctoria}" 327 "Ligules white; pappus absent" Anthemis 328 "All florets producing achenes, or only disc florets producing achenes" 329 328 "Only outer florets producing achenes" 341 329 "Only disc florets producing achenes" "Mauranthemum {Mauranthemum paludosum}" 329 "All florets producing achenes" 330 330 "Achenes heteromorphic, inner and outer achenes different" 331 330 "Achenes homomorphic" 337 331 "At least inner achenes curved to C-shape" Calendula 331 "All achenes straight or almost so" 332 332 "Annual herbs" 333 332 "Perennial herbs, without glands" 336 333 "Plants glandular-puberulous" 334 333 "Plants not glandular-puberulous (achenes dimorphic)" 335 334 "Achenes dimorphic, 2- and 3-angled" Dimorphotheca 334 "Achenes polymorphic, curved or straight, winged or wingless, sometimes rostrate" "Oligocarpus {Oligocarpus calendulaceus}" 335 "Achenes entirely lacking pappus" Glebionis 335 "Achenes with a very short scarious pappus, a solitary scale on ray achenes, a short ring on disc achenes" Matricaria 336 "Ray achenes shortly winged" "Argyranthemum {Argyranthemum frutescens}" 336 "Ray achenes ±terete" Leucanthemum 337 "Receptacle convex to conical" 338 337 "Receptacle flat" 340 338 "Ray ligules short, <10 mm long" "Asteraceae tribe Astereae" 338 "Ray ligules 10–18 mm long" 339 339 "Leaves 3-pinnatisect, at least at base; rays white" "Tripleurospermum {Tripleurospermum maritimum subsp. inodorum}" 339 "Leaves 1-pinnatisect; rays yellow" Euryops 340 "Achenes oblong to obovoid" Cymbonotus 340 "Achenes prominently winged" "Monoculus {Monoculus monstrosus}" 341 "Achenes with fleshy outer tissue" "Chrysanthemoides {Chrysanthemoides monilifera}" 341 "Achenes all dry" 342 342 "Achenes with prominent wings" Osteospermum 342 "Achenes without wings" 343 343 "Outer involucral bracts much longer than inner ones" "Asteraceae tribe Astereae" 343 "Involucral bracts c. equisized" 344 344 "Inner disc florets fertile" Cotula 344 "Inner disc florets functionally staminate" Leptinella