1 "Heads (i.e. whole inflorescence) forming burrs in fruit, or spiny and borne sessile in axils of leaves along stems." 2 1 "Heads not forming burrs in fruit or if with stiff barbed bristles then borne in terminal heads." 28 2 "Fruit (i.e. achenes plus fused involucral bracts) spiny or burrs, in axils of leaves along stems (flower heads sometimes in spike-like inflorescences)." 3 2 "Achenes not inside a fused fruit-like structure (sometimes enfolded by a conduplicate, unfused bract or scale); flower head not forming a burr, if spiny or with barbed hairs then borne in terminal heads." 5 3 "Leaves opposite, sometimes alternate above; fruit consisting of 1 ray achene enveloped by one of the modified inner involucral bracts; 1–several borne on the common receptacle, within the outer involucral bracts." Acanthospermum 3 "All leaves spirally arranged or in basal rosette; all the involucral bracts united to form a hard burr or nut enclosing 1 or more achenes; 1–several fruits borne in the leaf axils." 4 4 "Leaves entire, toothed, or if lobed then fruits covered with recurved spines." Xanthium 4 "Leaves deeply divided." Ambrosia 5 "Bracts and whole plant covered with glandular hairs." 6 5 "Bracts and whole plant without glands." 7 6 "Leaves linear, opposite below, alternate above." Madia 6 "Leaves lanceolate to elliptic, opposite." Sigesbeckia 7 "Corolla of outer florets longer than 8 mm." 8 7 "Corolla of outer florets up to 5 mm long." 18 8 "Leaves entire, toothed, or shallowly lobed." 9 8 "Leaves, at least lower ones, deeply lobed." 16 9 "Leaves sessile and usually clasping stem." 10 9 "Leaves petiolate or constricted towards base and not clasping stem." 11 10 "Leaves usually entire; involucral bracts in at least 3 rows." Zinnia 10 "Leaves usually toothed; involucral bracts in 2 rows." Guizotia 11 "Pappus of many hairs." 12 11 "Pappus of few hairs, absent, or otherwise." 13 12 "Plants prostrate." Tridax 12 "Plants erect shrubs or subshrubs." Olearia 13 "Achenes winged." Verbesina 13 "Achenes angled or compressed but without wings." 14 14 "Receptacle conical." Acmella 14 "Receptacle flat or convex." 15 15 "Heads in groups of 3; leaves ovate to rhombic, toothed." Melanthera 15 "Heads solitary; leaves narrow-lanceolate, more or less entire to toothed or minutely lobed." Wedelia 16 "Lobes of leaves up to 2 mm wide." Cosmos 16 "Lobes 4–50 mm wide." 17 17 "Shrubs 1–4 m high; ligules of florets white." Montanoa 17 "Subshrubs less than 1 m high; ligules of florets yellow." Coreopsis 18 "Receptacle naked." 19 18 "Receptacle with scales or chaffy." 20 19 "Involucral bracts in 1 row and fused; plant with pungent smell." Tagetes 19 "Involucral bracts in several rows, free." Schkuhria 20 "Pappus consisting of bristles or scales." 21 20 "Pappus absent, rarely a cup, very short hairs or scales." 22 21 "Leaves pinnately divided." Bidens 21 "Leaves entire or toothed." Galinsoga 22 "Bracts covered with stalked glands." Sigesbeckia 22 "Bracts without glands." 23 23 "Tubular florets sterile; involucral bracts in 1 row or if in 2 rows then outer row with 2–5 bracts." 24 23 "All florets fertile; involucral bracts numerous, in 2 or more rows." 26 24 "Leaves opposite; plants aquatic to semi-aquatic with soft creeping stems rooting at the nodes." Enydra 24 "Leaves alternate (lower ones often opposite) or clustered; lacking the above combination of features." 25 25 "Involucral bracts enfolding the ray achenes, with shortly spiny tips, sparsely hairy." Hemizonia 25 "Involucral bracts not enfolding the ray achenes, with membranous margins, woolly." Eriocephalus 26 "Receptacle conical." Acmella 26 "Receptacle flat or slightly convex." 27 27 "Outer achenes triangular in section, inner ones compressed." Eclipta 27 "All achenes in flower head similar." Wedelia 28 "Flower heads with all florets ligulate (not including petaloid stigmas that sometimes look like ligules); with milky latex." 29 28 "Flower heads with all florets tubular or some florets ligulate and some tubular; lacking milky latex." 52 29 "Florets blue or violet, rarely white." 30 29 "Florets yellow." 31 30 "Pappus of feathery hairs." Tragopogon 30 "Pappus of minute scales." Cichorium 31 "Receptacle of flower head with scales." 32 31 "Receptacle naked." 33 32 "Stems with spiny wings; plants spiny." Scolymus 32 "Stems without wings; soft herbs." Hypochaeris 33 "Involucral bracts in 1 whorl." 34 33 "Involucral bracts in more than 1 whorl and often with much smaller ones at base of flower head." 35 34 "Pappus feathery." Urospermum 34 "Pappus scaly or bristle-like." Hedypnois 35 "All leaves in basal rosette." 36 35 "Leaves basal and/or on erect stem (cauline leaves often much smaller)." 39 36 "All, or most achenes with a filiform beak longer than the fertile portion; scaly-muricate or muricate-spinulose in upper part." Taraxacum 36 "Achenes lacking filiform beak or beak not filiform and less than half the length of the fertile portion." 37 37 "Pappus of free scarious scales with acuminate apices produced into barbellate bristles." Microseris 37 "Pappus of numerous plumose or barbellate bristles." 38 38 "Bristles barbellate; pappus of inner and outer achenes similar." Youngia 38 "Bristles plumose; pappus of inner and outer achenes frequently dissimilar." Leontodon 39 "Achenes drawn into long neck." 40 39 "Achenes abruptly truncated at apex." 44 40 "Pappus hairs feathery." 41 40 "Pappus hairs simple." 42 41 "Achenes tapered at apex or more or less truncate; hairs 2-fid, base not expanded." Picris 41 "Achenes abruptly narrowed at apex into a filiform beak; hairs 3–5-fid, base expanded." Helminthotheca 42 "Achenes strongly compressed." Lactuca 42 "Achenes not compressed." 43 43 "Achenes with at least 1 ring of teeth below beak." Chondrilla 43 "Achenes without ring of teeth." Crepis 44 "All achenes laterally compressed." 45 44 "Achenes not laterally compressed, or sometimes the outer ones dorsiventrally compressed." 46 45 "Achenes elliptic in outline, both ends narrowed, sometimes cylindrical, margins wingless or narrow-winged; ribs never raised to form small wings." Sonchus 45 "Achenes flask-shaped in outline, constricted at the apex, margins with broad flat wings; ribs may be raised to form smaller secondary wings." Actites 46 "Pappus hairs feathery." 47 46 "Pappus hairs simple or absent." 48 47 "Achenes with a pale thickened hollow base c. a third as long as achene." Scorzonera 47 "Achenes not thickened at base." Picris 48 "Pappus absent, achenes with 20 ribs." Lapsana 48 "Pappus present, rarely absent and then achenes with 4 ribs." 49 49 "Achenes with 6 to several ribs." 50 49 "Achenes rough and 4-ribbed." 51 50 "Involucral bracts and upper peduncle with glandular hairs." Crepis 50 "Involucral bracts and upper peduncle glabrous." Youngia 51 "Pappus scale-like broadened at the base." Tolpis 51 "Pappus fine silky hairs." Reichardia 52 "All florets tubular and/or some filiform, or if some ligulate then ligules less than 3 mm long (involucral bracts sometimes coloured and resembling ligules or sometimes lobes of corolla long and resembling ligules)." 53 52 "Outer florets with ligules more than 3 mm long." 271 53 "Receptacle bearing inner tubular florets with scales or bristles; leaves usually spinose (thistles); styles with tuft of hairs below stigma branches." 54 53 "Receptacle glabrous or if with scales then leaves not spinose; styles glabrous." 65 54 "Flower heads with 1 floret, grouped into globose (compound) heads." Echinops 54 "Flower heads usually with many florets, not grouped into compound heads." 55 55 "Receptacle of flower head glabrous." Onopordum 55 "Receptacle with scales or bristles." 56 56 "Receptacle with scales." 57 56 "Receptacle with bristles." 59 57 "Achenes more or less cylindrical and ribbed." Hemisteptia 57 "Achenes compressed or flat." 58 58 "Involucral bracts with spreading spines." Cirsium 58 "Involucral bracts without spines and recurved or hooked." Arctium 59 "Involucral bracts loose and leaf-like." Carthamus 59 "Involucral bracts scale-like and more or less appressed." 60 60 "Achenes obliquely attached to receptacle." Centaurea 60 "Achenes vertically attached to receptacle." 61 61 "Leaves white-veined or variegated above; achenes compressed; anther filaments joined into a tube which encloses the basal appendages of the anthers." Silybum 61 "Leaves usually uniformly coloured above; achenes terete, sometimes ribbed; anther filaments free." 62 62 "Pappus 2 rows of bristles; the outer long and scabrous, the inner much shorter (less than half as long) and ciliate." Cnicus 62 "Pappus not composed of 2 rows of bristles of markedly differing length." 63 63 "Involucral bracts not ending in spines." Rhaponticum 63 "Involucral bracts ending in spines." 64 64 "Pappus hairs simple." Carduus 64 "Pappus hairs feathery." Cynara 65 "Leaves opposite, at least lower ones." 66 65 "Leaves alternately arranged and/or in basal rosette." 116 66 "Leaves opposite at least at base, rarely in whorls." 67 66 "Leaves alternately arranged and/or radical." 108 67 "Flower heads compound." 68 67 "Flower heads simple." 89 68 "Leaves opposite; dichotomously branching (repeatedly branching from below terminal inflorescence)." Flaveria 68 "Leaves all or mostly alternate, or crowded." 69 69 "Leaves 1–2-pinnatifid; margins spinose." Echinops 69 "Leaves with margins toothed or entire, margins not spinose." 70 70 "Branches with herbaceous wings extending from leaf bases down the internode; leaves mostly toothed." Pterocaulon 70 "Branches not winged; leaves entire." 71 71 "Common involucre surrounding the compound head of numerous scarious, opaque white or yellow bracts." Myriocephalus 71 "Common involucre of few green leaves, scales or absent." 72 72 "Heads with 2 types of florets (tubular and more or less filiform)." 73 72 "Heads with 1 type of floret." 75 73 "Leaves spathulate with petiole usually distinctly longer than the broad-circular to broad-obovate lamina." Stuartina 73 "Leaves linear, lanceolate, or oblanceolate." 74 74 "Plants with erect stems." Gnaphalium 74 "Plants stemless." Isoetopsis 75 "Inflorescence sessile at ground level, usually in a rosette of leaves." 76 75 "Inflorescence terminal on aerial stems or scapes." 77 76 "Pappus of 4 or 5 stiff bristles fused at the base." Actinobole 76 "Pappus absent." Chthonocephalus 77 "Pappus absent." 78 77 "Pappus present." 84 78 "Corolla hairy; common involucre absent; florets 20–40 per individual head." Eriochlamys 78 "Corolla lacking hairs; common involucre present; florets 1 or 2 per head." 79 79 "Involucral bracts papillate at tips; leaves hairy at the base only." Pogonolepis 79 "Involucral bracts not papillate at tips; leaves usually densely hairy." 80 80 "Pappus of 8–12 subplumose bristles, united at the base for about half the length of corolla tube. Plants prostrate, with reddish, somewhat wiry stems terminating in very woolly compound heads." Lemooria 80 "Pappus absent or not as above; plants lacking the above combination of features." 81 81 "General receptacle with long bristles." Trichanthodium 81 "General receptacle glabrous." 82 82 "Achenes with dense, entwined hairs." Gnephosis 82 "Achenes papillose, glabrous." 83 83 "Bracts subtending heads more or less similar to involucral bracts (except sometimes more concave); achenes brown." Angianthus 83 "Bracts subtending heads unlike involucral bracts; achenes pink or purple." Gnephosis 84 "Achenes tapering to long terminal beak." Waitzia 84 "Achenes otherwise." 85 85 "Pappus much shorter than the corolla." Gnephosis 85 "Pappus more or less equal in length to corolla." 86 86 "Achenes glabrous to papillose." Calocephalus 86 "Achenes hairy." 87 87 "Inflorescence branching, heads and branches subtended by large cordate leaves or bracts that ± enclose the head." Elephantopus 87 "Inflorescence simple, rarely branching from within the heads; heads with or without small enclosing bracts that in no way envelope the heads." 88 88 "Heads pedunculate within the compound head; bracts and pappus hairs white (or bracts brown-tinged)." Craspedia 88 "Heads sessile within the compound head; bracts and pappus hairs yellow." Pycnosorus 89 "Leaves lobed to deeply divided." 90 89 "Leaves entire or toothed." 94 90 "Pappus absent; water plant with soft branches; involucral bracts 4 and free." Enydra 90 "Pappus hair-like or with bristles or scales; not a water plant; involucral bracts more than 4 or fused." 91 91 "Upper leaves clasping stem; pappus of fine hairs." Emilia 91 "All leaves not clasping; pappus of few bristles or of scales." 92 92 "Involucral bracts fused into tube; plant with pungent smell." Tagetes 92 "Involucral bracts free; plants usually odourless." 93 93 "Pappus of 2–4 stiff, barbed awns; receptacle scales present, but not persistent." Bidens 93 "Pappus of 8–10 membranous scales (sometimes darkly striated at the apex); receptacle scales absent." Schkuhria 94 "Achenes narrow-cylindrical with a long straight or curved beak; outer florets strongly decurved and overhanging involucre." Millotia 94 "Achenes and florets lacking the above combination of characters." 95 95 "Pappus absent or very small." 96 95 "Pappus at least half as long as achenes or gland-tipped." 100 96 "Flower head covered with glandular hairs." 97 96 "Flower heads without glands." 98 97 "Style branches slightly exserted but not petaloid; ray florets with very small ligules; receptacle with somewhat persistent scales." Sigesbeckia 97 "Style branches long-exserted and petal-like; all florets tubular; receptacle naked." Gymnocoronis 98 "Achenes flat with involute herbaceous wings with large incurved appendages at the apex; small annual herbs." Ceratogyne 98 "Achenes lacking involute wings with apical appendages; large annual to perennial herbs." 99 99 "Receptacle naked (occasionally with innermost involucral bracts scale-like or scales sporadic); all florets tubular; inconspicuous pappus present." Ageratum 99 "Receptacle with scales; ray florets shortly ligulate; pappus absent." Enydra 100 "Involucral bracts fused at base and with glandular hairs; flower heads with 4 or 5 florets." Stevia 100 "Involucral bracts not as above; flower heads with numerous florets." 101 101 "Involucral bracts membranous." 102 101 "Involucral bracts herbaceous sometimes with membranous margins." 103 102 "All florets bisexual." Rutidosis 102 "Few outer florets female." Helichrysum 103 "Pappus hairs gland-tipped." Adenostemma 103 "Pappus without glands." 104 104 "Leaves linear to oblong, entire." 105 104 "Leaves lanceolate to ovate, toothed." 106 105 "Involucral bracts linear to linear-lanceolate." Glossocardia 105 "Involucral bracts broad." Acmella 106 "Pappus of 5 or more fragile bristles, soon falling from the achene." Ageratina 106 "Pappus of scales, awns or rudimentary." 107 107 "Pappus of scales, usually with awns." Ageratum 107 "Pappus 2–4 stiff, retrorsely barbed awns." Bidens 108 "Pappus purplish-mauve." Erechtites 108 "Pappus white or almost so." 109 109 "Florets orange to deep red or purple at least at top." 110 109 "Florets yellow to white." 111 110 "Leaves clasping upper stems." Emilia 110 "Leaves not clasping stems." Crassocephalum 111 "Marginal florets sterile." Arrhenechthites 111 "Marginal florets fertile." 112 112 "Achenes narrow-cylindrical with a long straight or curved beak; outer florets strongly decurved and overhanging involucre." Millotia 112 "Achenes and florets lacking the above combination of characters." 113 113 "Outer achenes flattened." Cineraria 113 "Outer achenes not flattened." 114 114 "Plants scandent semi-succulent twiners or trailers." Delairea 114 "Plants erect or decumbent, occasionally prostrate but never scandent, twining nor diffuse." 115 115 "Annual to perennial herbs, or shrubs with ligulate ray florets." Senecio 115 "Shrubs or small trees, florets all tubular." Bedfordia 116 "Involucral bracts in 1 whorl, rarely with a few much shorter ones at base; usually fused, although sometimes only at the base; flower heads simple." 117 116 "Involucral bracts in 2 or more whorls and free, or with compound flower heads." 167 117 "Leaves opposite at least at base, rarely in whorls." 118 117 "Leaves alternately arranged and/or radical." 159 118 "Flower heads compound." 119 118 "Flower heads simple." 140 119 "Leaves opposite; dichotomously branching (repeatedly branching from below terminal inflorescence)." Flaveria 119 "Leaves all or mostly alternate, or crowded." 120 120 "Leaves 1–2-pinnatifid; margins spinose." Echinops 120 "Leaves with margins toothed or entire, margins not spinose." 121 121 "Branches with herbaceous wings extending from leaf bases down the internode; leaves mostly toothed." Pterocaulon 121 "Branches not winged; leaves entire." 122 122 "Common involucre surrounding the compound head of numerous scarious, opaque white or yellow bracts." Myriocephalus 122 "Common involucre of few green leaves, scales or absent." 123 123 "Heads with 2 types of florets (tubular and more or less filiform)." 124 123 "Heads with 1 type of floret." 126 124 "Leaves spathulate with petiole usually distinctly longer than the broad-circular to broad-obovate lamina." Stuartina 124 "Leaves linear, lanceolate, or oblanceolate." 125 125 "Plants with erect stems." Gnaphalium 125 "Plants stemless." Isoetopsis 126 "Inflorescence sessile at ground level, usually in a rosette of leaves." 127 126 "Inflorescence terminal on aerial stems or scapes." 128 127 "Pappus of 4 or 5 stiff bristles fused at the base." Actinobole 127 "Pappus absent." Chthonocephalus 128 "Pappus absent." 129 128 "Pappus present." 135 129 "Corolla hairy; common involucre absent; florets 20–40 per individual head." Eriochlamys 129 "Corolla lacking hairs; common involucre present; florets 1 or 2 per head." 130 130 "Involucral bracts papillate at tips; leaves hairy at the base only." Pogonolepis 130 "Involucral bracts not papillate at tips; leaves usually densely hairy." 131 131 "Pappus of 8–12 subplumose bristles, united at the base for about half the length of corolla tube. Plants prostrate, with reddish, somewhat wiry stems terminating in very woolly compound heads." Lemooria 131 "Pappus absent or not as above; plants lacking the above combination of features." 132 132 "General receptacle with long bristles." Trichanthodium 132 "General receptacle glabrous." 133 133 "Achenes with dense, entwined hairs." Gnephosis 133 "Achenes papillose, glabrous." 134 134 "Bracts subtending heads more or less similar to involucral bracts (except sometimes more concave); achenes brown." Angianthus 134 "Bracts subtending heads unlike involucral bracts; achenes pink or purple." Gnephosis 135 "Achenes tapering to long terminal beak." Waitzia 135 "Achenes otherwise." 136 136 "Pappus much shorter than the corolla." Gnephosis 136 "Pappus more or less equal in length to corolla." 137 137 "Achenes glabrous to papillose." Calocephalus 137 "Achenes hairy." 138 138 "Inflorescence branching, heads and branches subtended by large cordate leaves or bracts that ± enclose the head." Elephantopus 138 "Inflorescence simple, rarely branching from within the heads; heads with or without small enclosing bracts that in no way envelope the heads." 139 139 "Heads pedunculate within the compound head; bracts and pappus hairs white (or bracts brown-tinged)." Craspedia 139 "Heads sessile within the compound head; bracts and pappus hairs yellow." Pycnosorus 140 "Leaves lobed to deeply divided." 141 140 "Leaves entire or toothed." 145 141 "Pappus absent; water plant with soft branches; involucral bracts 4 and free." Enydra 141 "Pappus hair-like or with bristles or scales; not a water plant; involucral bracts more than 4 or fused." 142 142 "Upper leaves clasping stem; pappus of fine hairs." Emilia 142 "All leaves not clasping; pappus of few bristles or of scales." 143 143 "Involucral bracts fused into tube; plant with pungent smell." Tagetes 143 "Involucral bracts free; plants usually odourless." 144 144 "Pappus of 2–4 stiff, barbed awns; receptacle scales present, but not persistent." Bidens 144 "Pappus of 8–10 membranous scales (sometimes darkly striated at the apex); receptacle scales absent." Schkuhria 145 "Achenes narrow-cylindrical with a long straight or curved beak; outer florets strongly decurved and overhanging involucre." Millotia 145 "Achenes and florets lacking the above combination of characters." 146 146 "Pappus absent or very small." 147 146 "Pappus at least half as long as achenes or gland-tipped." 151 147 "Flower head covered with glandular hairs." 148 147 "Flower heads without glands." 149 148 "Style branches slightly exserted but not petaloid; ray florets with very small ligules; receptacle with somewhat persistent scales." Sigesbeckia 148 "Style branches long-exserted and petal-like; all florets tubular; receptacle naked." Gymnocoronis 149 "Achenes flat with involute herbaceous wings with large incurved appendages at the apex; small annual herbs." Ceratogyne 149 "Achenes lacking involute wings with apical appendages; large annual to perennial herbs." 150 150 "Receptacle naked (occasionally with innermost involucral bracts scale-like or scales sporadic); all florets tubular; inconspicuous pappus present." Ageratum 150 "Receptacle with scales; ray florets shortly ligulate; pappus absent." Enydra 151 "Involucral bracts fused at base and with glandular hairs; flower heads with 4 or 5 florets." Stevia 151 "Involucral bracts not as above; flower heads with numerous florets." 152 152 "Involucral bracts membranous." 153 152 "Involucral bracts herbaceous sometimes with membranous margins." 154 153 "All florets bisexual." Rutidosis 153 "Few outer florets female." Helichrysum 154 "Pappus hairs gland-tipped." Adenostemma 154 "Pappus without glands." 155 155 "Leaves linear to oblong, entire." 156 155 "Leaves lanceolate to ovate, toothed." 157 156 "Involucral bracts linear to linear-lanceolate." Glossocardia 156 "Involucral bracts broad." Acmella 157 "Pappus of 5 or more fragile bristles, soon falling from the achene." Ageratina 157 "Pappus of scales, awns or rudimentary." 158 158 "Pappus of scales, usually with awns." Ageratum 158 "Pappus 2–4 stiff, retrorsely barbed awns." Bidens 159 "Pappus purplish-mauve." Erechtites 159 "Pappus white or almost so." 160 160 "Florets orange to deep red or purple at least at top." 161 160 "Florets yellow to white." 162 161 "Leaves clasping upper stems." Emilia 161 "Leaves not clasping stems." Crassocephalum 162 "Marginal florets sterile." Arrhenechthites 162 "Marginal florets fertile." 163 163 "Achenes narrow-cylindrical with a long straight or curved beak; outer florets strongly decurved and overhanging involucre." Millotia 163 "Achenes and florets lacking the above combination of characters." 164 164 "Outer achenes flattened." Cineraria 164 "Outer achenes not flattened." 165 165 "Plants scandent semi-succulent twiners or trailers." Delairea 165 "Plants erect or decumbent, occasionally prostrate but never scandent, twining nor diffuse." 166 166 "Annual to perennial herbs, or shrubs with ligulate ray florets." Senecio 166 "Shrubs or small trees, florets all tubular." Bedfordia 167 "Flower heads clustered into compound heads with scales or leaves more or less densely arranged around it, rarely without." 168 167 "Flower heads solitary and/or stalked." 189 168 "Leaves opposite; dichotomously branching (repeatedly branching from below terminal inflorescence)." Flaveria 168 "Leaves all or mostly alternate, or crowded." 169 169 "Leaves 1–2-pinnatifid; margins spinose." Echinops 169 "Leaves with margins toothed or entire, margins not spinose." 170 170 "Branches with herbaceous wings extending from leaf bases down the internode; leaves mostly toothed." Pterocaulon 170 "Branches not winged; leaves entire." 171 171 "Common involucre surrounding the compound head of numerous scarious, opaque white or yellow bracts." Myriocephalus 171 "Common involucre of few green leaves, scales or absent." 172 172 "Heads with 2 types of florets (tubular and more or less filiform)." 173 172 "Heads with 1 type of floret." 175 173 "Leaves spathulate with petiole usually distinctly longer than the broad-circular to broad-obovate lamina." Stuartina 173 "Leaves linear, lanceolate, or oblanceolate." 174 174 "Plants with erect stems." Gnaphalium 174 "Plants stemless." Isoetopsis 175 "Inflorescence sessile at ground level, usually in a rosette of leaves." 176 175 "Inflorescence terminal on aerial stems or scapes." 177 176 "Pappus of 4 or 5 stiff bristles fused at the base." Actinobole 176 "Pappus absent." Chthonocephalus 177 "Pappus absent." 178 177 "Pappus present." 184 178 "Corolla hairy; common involucre absent; florets 20–40 per individual head." Eriochlamys 178 "Corolla lacking hairs; common involucre present; florets 1 or 2 per head." 179 179 "Involucral bracts papillate at tips; leaves hairy at the base only." Pogonolepis 179 "Involucral bracts not papillate at tips; leaves usually densely hairy." 180 180 "Pappus of 8–12 subplumose bristles, united at the base for about half the length of corolla tube. Plants prostrate, with reddish, somewhat wiry stems terminating in very woolly compound heads." Lemooria 180 "Pappus absent or not as above; plants lacking the above combination of features." 181 181 "General receptacle with long bristles." Trichanthodium 181 "General receptacle glabrous." 182 182 "Achenes with dense, entwined hairs." Gnephosis 182 "Achenes papillose, glabrous." 183 183 "Bracts subtending heads more or less similar to involucral bracts (except sometimes more concave); achenes brown." Angianthus 183 "Bracts subtending heads unlike involucral bracts; achenes pink or purple." Gnephosis 184 "Achenes tapering to long terminal beak." Waitzia 184 "Achenes otherwise." 185 185 "Pappus much shorter than the corolla." Gnephosis 185 "Pappus more or less equal in length to corolla." 186 186 "Achenes glabrous to papillose." Calocephalus 186 "Achenes hairy." 187 187 "Inflorescence branching, heads and branches subtended by large cordate leaves or bracts that ± enclose the head." Elephantopus 187 "Inflorescence simple, rarely branching from within the heads; heads with or without small enclosing bracts that in no way envelope the heads." 188 188 "Heads pedunculate within the compound head; bracts and pappus hairs white (or bracts brown-tinged)." Craspedia 188 "Heads sessile within the compound head; bracts and pappus hairs yellow." Pycnosorus 189 "Pappus absent or reduced to a vestigial scarious cup." 190 189 "Pappus of bristles or scales at least subequal to the achene." 219 190 "Leaves entire or toothed." 191 190 "Leaves deeply divided and pinnatifid." 213 191 "Achenes drawn into a terminal beak." 192 191 "Achenes more or less abruptly truncated at apex." 195 192 "Rosette forming perennial." 193 192 "Erect or low-growing annual, biennial or perennial." 194 193 "Ray florets with small but well-developed ligules; beak of achenes well-developed, often densely glandular, apex with a thickened pale collar." Lagenophora 193 "Ray florets with ligules rudimentary or absent; beak of achenes rudimentary or absent, glands lacking or rarely with a few near the apex, thickened apical collar absent." Solenogyne 194 "Achenes with short beaks (less than a quarter the length of achene), glandular near apex; erect herb 30–60 cm high, flower heads nodding." Carpesium 194 "Achenes with long beaks (more than half the length of achene), woolly near apex; small more or less prostrate herb with stems 1–4 cm long, flower heads erect." Millotia 195 "Florets blue to purple." 196 195 "Florets yellow or white to light green, sometimes tinged red." 197 196 "Achenes usually 4-angled or ribbed; glandular, with thickened apical ring resembling pappus." Ethulia 196 "Achenes 8 to 10 ribs; non-glandular, pappus of bristles (often caducous)." Centratherum 197 "Receptacle with scales; flower head usually surrounded by numerous white, papery involucral bracts 5–10 mm long." Ammobium 197 "Receptacle naked; flower head, if surrounded by white involucral bracts, with a pappus of non-persistent bristles." 198 198 "All florets tubular to campanulate." 199 198 "Outer florets reduced, filiform or with a small ligule; differing in shape from the inner florets." 205 199 "Achenes dark brown to black, tuberculate; herb to c. 60 cm high, more or less woolly." Acomis 199 "Achenes not dark brown to black, nor tuberculate; habit various." 200 200 "Involucral bracts multiseriate." 201 200 "Involucral bracts in 2 rows." 202 201 "Shrubs; glabrous to minutely glandular hairy, viscid; involucral bracts pale golden brown to straw-coloured." Haeckeria 201 "Biennial herbs; more or less woolly, aromatic; involucral bracts pale brown tinged with pink, occasionally almost colourless." Calomeria 202 "Annual herbs with wiry stems; corolla woolly at base, glandular near apex." Eriochlamys 202 "Annual or perennial herbs lacking wiry stems; corolla glabrous and non-glandular." 203 203 "Leaves lyrate-pinnatifid; outer florets tubular, inner ones campanulate." Dichrocephala 203 "Leaves entire to pinnate; florets tubular." 204 204 "Achenes pedicellate, dimorphic; outer glandular-hairy, winged; inner sparsely glandular-hairy, with narrow marginal ridge." Cotula 204 "Achenes non-pedicellate; outer glabrous, not winged; inner failing to develop." Abrotanella 205 "Pappus of bristle-like hairs present when flowering, caducous at achenes maturity." 206 205 "Pappus absent or minute throughout floral development." 207 206 "Achenes sparsely to densely hairy; involucral bracts never clawed, herbaceous with scarious margins." Blumea 206 "Achenes glabrous; involucral bracts with scarious claw, inner bracts with white or coloured lamina." Hyalosperma 207 "Flower heads sessile; achenes with spines (2 ciliate horns) at apex." Ceratogyne 207 "Flower heads on a distinct stalk; if sessile, achenes without horns." 208 208 "All leaves in a basal rosette." 209 208 "Leaves along erect or creeping stems and at base." 210 209 "Ray florets with small but well-developed ligules; beak of achenes well-developed, often densely glandular, apex with a thickened pale collar." Lagenophora 209 "Ray florets with ligules rudimentary or absent; beak of achenes rudimentary or absent, glands lacking or rarely with a few near the apex, thickened apical collar absent." Solenogyne 210 "Flower heads solitary on terminal peduncles." Cotula 210 "Flower heads either in terminal panicles, racemes or axillary and almost sessile." 211 211 "Ray florets with small, deeply 3-lobed, yellow ligules." Madia 211 "Ray florets tubular to filiform." 212 212 "Achenes with 3–5 obtuse ribs which frequently fail to extend to the apex, usually pubescent." Centipeda 212 "Achenes with 8–10 fine ribs reaching to the apex, glabrous except for a few bristles at the base." Epaltes 213 "Shrub with persistent woody stems." Artemisia 213 "Annual or perennial herbs, rarely subshrubs with a short woody base." 214 214 "Flower heads sessile among the leaves; achenes with 1 or more apical spines." Soliva 214 "Flower heads pedunculate; achenes spineless." 215 215 "Flower heads numerous in dense compound corymbs or large terminal panicles." 216 215 "Flower heads solitary or in corymbs of 2 or 3." 217 216 "Receptacle scales absent." Tanacetum 216 "Receptacle scales present." Parthenium 217 "Florets sessile on the receptacle, which appears slightly pitted when the achenes have fallen; plants erect to ascending." Matricaria 217 "Florets raised above the receptacle on minute pedicels which persist when the achenes have fallen, plants erect; or florets sessile, plants creeping." 218 218 "Disc florets usually fertile, style bifid; achenes with blunt uniseriate hairs on surface; plants usually erect or decumbent." Cotula 218 "Disc florets sterile, style ending in circular disc; achenes glabrous; plants usually prostrate creeping herbs rooting at the nodes." Leptinella 219 "Plants dioecious (i.e. individual plants with flower heads containing either male or female florets)." 220 219 "Plants monoecious (i.e. flower heads usually with some bisexual flowers or male and female flowers on same plant)." 223 220 "Plants low-growing soft herbs." 221 220 "Plants shrubs or undershrubs with persistent woody stems." 222 221 "Achenes glabrous, dimorphic; pappus persistent; outer involucral bracts scarious." Parantennaria 221 "Achenes with appressed hairs; pappus not persistent; outer involucral bracts papery white at tip." Ewartia 222 "Flower heads terminal, solitary; leaves sessile." Cratystylis 222 "Flower heads in panicles; leaves petiolate." Baccharis 223 "Leaves crowded, spathulate, indented at apex and clasping stem." Facelis 223 "Leaves not as above." 224 224 "Pappus a cup or scales; margin of scales sometimes hair-like and feathery." 225 224 "Pappus of simple, barbed or feathery bristles." 229 225 "Receptacle scales present." Ammobium 225 "Receptacle scales absent." 226 226 "Achenes densely hairy." 227 226 "Achenes glabrous or with short globose hairs." 228 227 "Tufted herb; flower heads basal, sessile, densely crowded amongst long linear leaves." Isoetopsis 227 "Decumbent to ascending herb; flower heads terminal, pedunculate, overtopping short linear leaves." Elachanthus 228 "Leaves deeply dissected to pinnate; ray floret ligulate, solitary, yellow; pappus scales darkly striated at apex." Schkuhria 228 "Leaves entire; ray florets absent; pappus scales not darkly striated at apex." Rutidosis 229 "Shrubs with persistent woody stems." 230 229 "Annual or perennial herbs, rarely subshrubs with a short woody base." 234 230 "Branches spine-tipped." Acanthocladium 230 "Branches not spine-tipped." 231 231 "Flower heads with outer fertile female florets surrounding bisexual or functionally male florets." 232 231 "Flower heads with all florets similar." 233 232 "Female florets uniseriate; inner florets fertile, producing achenes." Olearia 232 "Female florets multiseriate; inner florets functionally male, not producing achenes." Pluchea 233 "Tips of involucral bracts straight to incurved; receptacle with scales." Cassinia 233 "Tips of involucral bracts spreading; receptacle lacking scales." Ozothamnus 234 "Ray florets with small ligules." 235 234 "Ray florets absent or lacking ligules." 244 235 "Outer involucral bracts sessile; inner bracts with scarious lamina on longer more or less herbaceous to hardened claws." Podolepis 235 "Outer involucral bracts not differing markedly from the inner ones; inner bracts lacking claws." 236 236 "Ligules and disc florets all yellow." 237 236 "Ligules white to blue, purple or pink; disc florets usually a contrasting colour, typically yellow or violet." 239 237 "Dwarf shrub with terete more or less succulent leaves." Kippistia 237 "Herbs with flat non-succulent leaves." 238 238 "Annual, sticky and aromatic due to glandular hairs; anthers tailed at the base." Solidago 238 "Perennial with rhizomes, non-glandular; anthers obtuse at the base." Dittrichia 239 "Achenes with long neck." Ixiochlamys 239 "Achenes lacking a long neck." 240 240 "Disc florets sterile, producing abortive achenes." Minuria 240 "Disc florets fertile." 241 241 "Involucral bracts a dark purple-brown, with white margins." Dichromochlamys 241 "Involucral bracts green." 242 242 "Flower heads numerous in panicles." Conyza 242 "Flower heads solitary or in loose groups of 2–4." 243 243 "Involucral bracts reflexed in fruit; disc florets yellow; achenes flattened." Erigeron 243 "Involucral bracts not reflexed in fruit; disc florets purple; achenes terete." Streptoglossa 244 "Leaves serrate to dissected or lobed." 245 244 "Leaves entire." 249 245 "Inner involucral bracts with fringed apices, far exceeded by the florets." Rhaponticum 245 "Inner involucral bracts with entire apices, subequal to the florets." 246 246 "Fertile achenes lacking a pappus of bristles (aborted achenes sometimes with 2–5 weak barbellate bristles)." Epaltes 246 "Achenes with a pappus of free bristles." 247 247 "Inner florets functionally male; achenes sterile." Pluchea 247 "Inner florets bisexual or functionally female, capable of producing fertile achenes." 248 248 "Pappus uniseriate." Blumea 248 "Pappus 2-seriate; outer bristles shorter than the inner ones." Vernonia 249 "Involucral bracts (at least the outer ones) with ciliate margin." 250 249 "Involucral bracts lacking a ciliate margin (but outer ones sometimes woolly or lacerate)." 253 250 "Lamina of the involucral bracts conspicuous, petaloid, papery, white to coloured; achenes with a distinct filiform, sterile beak." Waitzia 250 "Lamina of the involucral bracts inconspicuous, scarious to plumose, if conspicuously petaloid and white or coloured then achenes lacking beak." 251 251 "Flower heads terminal on short branches, sessile; usually surrounded by a cluster of leaves." Triptilodiscus 251 "Flower heads terminal on long peduncles; not surrounded by a cluster of leaves." 252 252 "Corollas overhanging the margin of the involucre; achenes 3- or 4-ribbed, often with a distinct sterile beak." Leptorhynchos 252 "Corollas not overhanging the involucre; achenes without ribs and lacking a beak." Chrysocephalum 253 "Involucral bracts uniseriate or almost so, equal, herbaceous." Millotia 253 "Involucral bracts multiseriate, frequently with inner bracts longer then the outer ones, herbaceous to scarious or coloured." 254 254 "Involucral bracts with lamina never petaloid, white or brightly coloured." 255 254 "Involucral bracts with lamina petaloid, papery, white to brightly coloured." 263 255 "All florets filiform." Scorzonera 255 "Florets tubular and filiform or all tubular." 256 256 "Outer filiform florets in 2 to several whorls." 257 256 "Filiform florets few to absent." 258 257 "Flower heads sessile to subsessile in dense globose to spicate clusters, frequently subtended by a whorl of leaves; involucral bracts scarious; plants woolly, especially lower leaf surface." Gnaphalium 257 "Flower heads pedunculate, usually in large panicles, not subtended by a whorl of leaves; involucral bracts herbaceous, sometimes with scarious margin; plants not woolly." Conyza 258 "Creeping, aromatic herb with glandular hairs; achenes on short pedicels which persist on the receptacle; florets tubular." Podotheca 258 "Habit not creeping, lacking the above combination of characters." 259 259 "Receptacle with scales; plant a white-tomentose aromatic herb with amplexicaul leaves." Apalochlamys 259 "Receptacle lacking scales; plant lacking the above combination of characters." 260 260 "Pappus 2-seriate; outer bristles shorter than the inner; not fused." Vernonia 260 "Pappus uniseriate; shortly fused at the base." 261 261 "Achenes with a slender sterile beak below the pappus." Leptorhynchos 261 "Achenes without a slender sterile beak below the pappus." 262 262 "Outer bracts often extending down the peduncle, broad and membranous, the herbaceous base reduced and broadly triangular." Leptorhynchos 262 "Outer bracts never extending down the peduncle, linear or oblong, green and herbaceous throughout except for membranous apex." Leiocarpa 263 "Outer and inner involucral bracts uniformly scarious, never clawed; flower heads yellow." Pseudognaphalium 263 "Inner and intermediate involucral bracts divided into a cartilaginous-herbaceous basal claw and a conspicuous lamina; colour of bracts various." 264 264 "Pappus plumose from the base (broader basal region sometimes lacks ornamentation)." 265 264 "Pappus either barbellate from the base, or plumose only at apex, or more or less subplumose at the base and barbellate at apex." 267 265 "Achenes hairy." Rhodanthe 265 "Achenes glabrous." 266 266 "Achenes ellipsoid, verrucose; pappus persistent, eventually breaking shortly above base; leaves with more or less flat lamina." Leucochrysum 266 "Achenes oblong to obovoid, smooth or papillose; pappus united in a ring at the base and falling as a unit; leaves more or less terete." Hyalosperma 267 "Florets purple, at least apically." Vellereophyton 267 "Florets yellow." 268 268 "Inflorescence a dense terminal cluster of small subsessile heads; female florets more numerous than the disc florets." Pseudognaphalium 268 "Inflorescence not as above; female florets less numerous than the disc florets." 269 269 "Pappus bristles shortly plumose at the apex." Anemocarpa 269 "Pappus bristles barbellate." 270 270 "Achenes prismatic, glabrous with papus yellow throughout; flower heads large, with many florets; involucral bracts numerous, all brightly coloured." Xerochrysum 270 "Achenes lacking the above combination of characters; flower heads usually smaller with few florets; outer involucral bracts hyaline, brownish or white." Helichrysum 271 "Leaves opposite, at least lower ones." 272 271 "Leaves alternately arranged and/or in basal rosette." 298 272 "Fruit (i.e. achenes plus fused involucral bracts) spiny or burrs, in axils of leaves along stems (flower heads sometimes in spike-like inflorescences)." 273 272 "Achenes not inside a fused fruit-like structure (sometimes enfolded by a conduplicate, unfused bract or scale); flower head not forming a burr, if spiny or with barbed hairs then borne in terminal heads." 275 273 "Leaves opposite, sometimes alternate above; fruit consisting of 1 ray achene enveloped by one of the modified inner involucral bracts; 1–several borne on the common receptacle, within the outer involucral bracts." Acanthospermum 273 "All leaves spirally arranged or in basal rosette; all the involucral bracts united to form a hard burr or nut enclosing 1 or more achenes; 1–several fruits borne in the leaf axils." 274 274 "Leaves entire, toothed, or if lobed then fruits covered with recurved spines." Xanthium 274 "Leaves deeply divided." Ambrosia 275 "Bracts and whole plant covered with glandular hairs." 276 275 "Bracts and whole plant without glands." 277 276 "Leaves linear, opposite below, alternate above." Madia 276 "Leaves lanceolate to elliptic, opposite." Sigesbeckia 277 "Corolla of outer florets longer than 8 mm." 278 277 "Corolla of outer florets up to 5 mm long." 288 278 "Leaves entire, toothed, or shallowly lobed." 279 278 "Leaves, at least lower ones, deeply lobed." 286 279 "Leaves sessile and usually clasping stem." 280 279 "Leaves petiolate or constricted towards base and not clasping stem." 281 280 "Leaves usually entire; involucral bracts in at least 3 rows." Zinnia 280 "Leaves usually toothed; involucral bracts in 2 rows." Guizotia 281 "Pappus of many hairs." 282 281 "Pappus of few hairs, absent, or otherwise." 283 282 "Plants prostrate." Tridax 282 "Plants erect shrubs or subshrubs." Olearia 283 "Achenes winged." Verbesina 283 "Achenes angled or compressed but without wings." 284 284 "Receptacle conical." Acmella 284 "Receptacle flat or convex." 285 285 "Heads in groups of 3; leaves ovate to rhombic, toothed." Melanthera 285 "Heads solitary; leaves narrow-lanceolate, more or less entire to toothed or minutely lobed." Wedelia 286 "Lobes of leaves up to 2 mm wide." Cosmos 286 "Lobes 4–50 mm wide." 287 287 "Shrubs 1–4 m high; ligules of florets white." Montanoa 287 "Subshrubs less than 1 m high; ligules of florets yellow." Coreopsis 288 "Receptacle naked." 289 288 "Receptacle with scales or chaffy." 290 289 "Involucral bracts in 1 row and fused; plant with pungent smell." Tagetes 289 "Involucral bracts in several rows, free." Schkuhria 290 "Pappus consisting of bristles or scales." 291 290 "Pappus absent, rarely a cup, very short hairs or scales." 292 291 "Leaves pinnately divided." Bidens 291 "Leaves entire or toothed." Galinsoga 292 "Bracts covered with stalked glands." Sigesbeckia 292 "Bracts without glands." 293 293 "Tubular florets sterile; involucral bracts in 1 row or if in 2 rows then outer row with 2–5 bracts." 294 293 "All florets fertile; involucral bracts numerous, in 2 or more rows." 296 294 "Leaves opposite; plants aquatic to semi-aquatic with soft creeping stems rooting at the nodes." Enydra 294 "Leaves alternate (lower ones often opposite) or clustered; lacking the above combination of features." 295 295 "Involucral bracts enfolding the ray achenes, with shortly spiny tips, sparsely hairy." Hemizonia 295 "Involucral bracts not enfolding the ray achenes, with membranous margins, woolly." Eriocephalus 296 "Receptacle conical." Acmella 296 "Receptacle flat or slightly convex." 297 297 "Outer achenes triangular in section, inner ones compressed." Eclipta 297 "All achenes in flower head similar." Wedelia 298 "Pappus present and usually prominent." 299 298 "Pappus absent, reduced to less than 1 mm or cup-like." 332 299 "Involucral bracts in 1 whorl, rarely with a few much shorter bracteoles at base; often slightly fused at base." 300 299 "Involucral bracts in 2 to several whorls, usually free." 303 300 "Pappus scale-like; heads solitary on long scape." Gazania 300 "Pappus of hairs; heads usually in terminal clusters, rarely solitary on long scapes." 301 301 "Leaves palmately veined; woody shrubs." Roldana 301 "Plants lacking the above combination of characters." 302 302 "Achenes not compressed." Senecio 302 "Achenes, at least those of the ray florets, compressed." Cineraria 303 "Receptacle with scales." 304 303 "Receptacle rough or pitted but without scales." 308 304 "Plants usually more than 1 m high and with 1 to several erect branches." 305 304 "Plants with basal rootstock, somewhat branched and with branches shorter than 50 cm long." 307 305 "Leaves deeply divided." Tithonia 305 "Leaves entire or toothed." 306 306 "Pappus of short bristles; apex of ligule of florets lobed or distinctly toothed." Verbesina 306 "Pappus of non-persistent scales; apex of ligule of florets scarcely toothed." Helianthus 307 "Ligules of ray florets scarcely longer than involucral bracts." Glossogyne 307 "Ligules large and spectacular, much longer than involucral bracts." Gaillardia 308 "Involucral bracts fused into tube." Gazania 308 "Involucral bracts in several rows, free." 309 309 "Leaves, at least some of them, divided or lobed." 310 309 "Leaves entire or toothed." 313 310 "Ray florets female, fertile." 311 310 "Ray florets sterile." 312 311 "Achenes with 3 wing-like ridges on one side; pappus of pinkish scales." Arctotis 311 "Achenes variously ribbed but lacking wing-like ridges; pappus of white to brownish bristles." Vittadinia 312 "Pappus double; outer obovate scales, inner bristles." Ursinia 312 "Pappus of small scales." Arctotheca 313 "Inner involucral bracts dry and membranous, with or without an herbaceous base." 314 313 "All involucral bracts herbaceous or sometimes with dry membranous margin." 315 314 "Achenes abruptly constricted below a short neck below the broader apex." Dittrichia 314 "Achenes ellipsoid to cylindrical." Podolepis 315 "Ray florets in 2 or more rows." 316 315 "Ray florets in 1 row." 325 316 "Achenes with a long neck." Ixiochlamys 316 "Achenes lacking a long neck." 317 317 "Tubular florets sterile; sterile achenes with pappus of long free barbellate bristles and shorter bristles often fused into scales or a cup." Minuria 317 "All florets fertile (but not always, producing achenes); pappus not as above." 318 318 "Involucral bracts of fruiting flower heads clasping; dark purple brown with white margins." Dichromochlamys 318 "Involucral bracts of fruiting flower heads spreading or recurved; herbaceous, with or without a scarious margin." 319 319 "Ligules absent or inconspicuous i.e. more or less as long as pappus; flower heads numerous, in panicles." 320 319 "Ligules exceeding the pappus; flower heads usually solitary or in loose groups of 2–4." 321 320 "Plant glabrous; involucral bracts tinged purple at apex and margins." Aster 320 "Plant pubescent to scabrous, at least on the upper stem; if glabrous then with a purple spot at the apex of the involucral bracts." Conyza 321 "Corolla of disc florets as long as or longer than the pappus, persisting." Camptacra 321 "Corolla of disc florets usually as long as or shorter than the pappus, not persisting." 322 322 "Plant glabrous." Aster 322 "Plant pubescent to scabrous." 323 323 "Achenes variously hairy but always with a basal tuft of hairs; base of corolla swollen around swollen style base; style branches flattened, with triangular appendages." Erigeron 323 "Achenes variously hairy but lacking a basal tuft of hairs; base of corolla not swollen; style branches not as above." 324 324 "Receptacle convex; achenes always with a basal tuft of hairs, bristles free." Vittadinia 324 "Receptacle flat; achenes hairy but lacking a basal tuft of hairs, bristles joined at the base." Streptoglossa 325 "Pappus double row of bristles; inner long, barbellate; outer short and inconspicuous; brick red to rust coloured." Heterotheca 325 "Pappus lacking the above combination of characters." 326 326 "Plants more or less resinous; involucral bracts 4–8-seriate, graduated, more or less indurated, middle and outer with spreading green tips." Grindelia 326 "Plants lacking the above combination of characters." 327 327 "Pappus of scales and/or awns or absent." 328 327 "Pappus 1 or 2 rows of barbellate bristles." 329 328 "Pappus of rigid, usually barbed awns, often alternating with scales, occasionally an entire scarious collar, or rarely absent; achene faces sometimes hairy but not enveloped in wool; plants not fleshy, leaves variously hairy but not tomentose." Calotis 328 "Pappus of scales; achenes enveloped in wool; plants fleshy, leaves tomentose, especially on lower surface." Arctotheca 329 "Pappus less than a quarter as long as the achenes." Brachyscome 329 "Pappus about as long as achenes or longer." 330 330 "Herbs with a basal rosette." Celmisia 330 "Shrubs and subshrubs; if herbaceous, lacking a basal rosette." 331 331 "Perennial with seasonal stems arising from a woody rootstock; corolla of disc florets persisting on mature achenes; anther appendages lanceolate, with central strand of darker tissue." Camptacra 331 "Shrubs and undershrubs; corolla of disc florets not persisting on mature achenes; anther appendages acute, lacking central strand of darker tissue." Olearia 332 "Achenes drawn into short beak." Lagenophora 332 "Achenes abruptly truncated at apex." 333 333 "Receptacle with scales." 334 333 "Receptacles naked." 337 334 "Flower heads small and in terminal clusters." Achillea 334 "Flower heads larger and individually stalked or solitary." 335 335 "Leaves lobed to almost entire." Cymbonotus 335 "Leaves deeply divided." 336 336 "Receptacle scales longer than outer florets, becoming hardened with age." Erodiophyllum 336 "Receptacle scales shorter than florets." Anthemis 337 "All florets producing achenes." 338 337 "Only outer florets producing achenes." 347 338 "Achenes from outer and inner florets different." 339 338 "Achenes all alike." 342 339 "At least inner achenes curved to C-shaped." Calendula 339 "All achenes straight or almost so." 340 340 "Annuals covered with glandular hairs." Dimorphotheca 340 "Perennials without glandular hairs." 341 341 "Ligules pale to golden yellow, rarely white but then usually yellow toward base." Chrysanthemum 341 "Ligules completely white." Leucanthemum 342 "Receptacle flat, ray florets yellow; leaves grey-white, tomentose beneath, less hairy and more or less green above." Cymbonotus 342 "Receptacle convex to conical, ray florets various; leaves concolorous, green." 343 343 "Achenes triquetrous; inner surface with 3 ribs; outer surface wrinkled, with 2 glands at apex; pappus a small crenulate corona; leaves mostly 3-pinnatisect, feathery in appearance." Tripleurospermum 343 "Achenes lacking the above combination of characters; leaves various." 344 344 "Plants aromatic, with chamomile-like odour; involucral bracts thinly scarious with expanded obtuse apices." 345 344 "Plants sometimes with sparse glands but never aromatic; involucral bracts usually with narrow scarious hyaline margins." 346 345 "Perennial with rhizomes; flower heads in dense compound corymbs." Tanacetum 345 "Annual; flower heads in loose cymes." Matricaria 346 "Plant a shrub; involucral bracts in 1 row, fused below; ligules yellow." Euryops 346 "Plant lacking the above combination of characters." Brachyscome 347 "Achenes with fleshy outer tissue." Chrysanthemoides 347 "Achenes all dry." 348 348 "Achenes with prominent wings." Osteospermum 348 "Achenes without wings" Grindelia