1 "Leaflets with a distinct articulation (not coloured swelling) at the base; cone scales in vertical rows" Zamia 1 "Leaflets not articulated at the base; cone scales in spiral arrangement" 2 2 "Leaflets not or hardly narrowed at the base; seeds on short stalks" Dioon 2 "Leaflets clearly narrowed at the base; seeds not on stalks" 3 3 "Cone scales with flattened 4 or 6-sided blunt ends; leaflets usually toothed" Encephalartos 3 "Cone scales pointed or with a terminal spine; leaflets without toothed margins" 4 4 "Leaflets joined to upper edge of midrib, not swollen or coloured at base; cones not or hardly stalked, scales without extended spines" Lepidozamia 4 "Leaflets attached at midline of the midrib, swollen and coloured at the base; cones stalked, scales with erect spines" Macrozamia